®ljf i. 1. Hill library 5fnrth (Harolina ^tatp (Ttillpgp . QK635 .y? :e'hiicnm , watery white to greenish. „ leucopus opaque w^iite. Reticularia lycoperdon ... white. Spumaria alba w^hite. IStemoidtis ferniginea , lemon yellow. „ fnsca ... . white. Tilmadoche mutablUs yellow. - Trichia affinis pure white. „ fallax white and rose. „ rnria white. Ann. Bat., Vol. IV. No. xiv. (1890). A Monogra'ph of the Mi/xogastres. 21 The Plasmodium of most species inliabitiiig dead leaves is discoloured by foreign matter before the change to sporangia takes place. The foregoing remarks may be summarized as follows — 1. The Myxogastres, as stated by De Bary, appear to have originated from a primitive group of aquatic organisms, the Flagellatae. 2. The sequence of differentiation which gives individuality to the group, is almost entirely confined to the reproductive phase, and follows in many important points, the lines of development observable in. the Fungi. 3. The subordinate position occupied by the Fungi and the Myxogastres in the general evolution of the vegetable kingdom may be traced to the absence of chromatophores. The lichen- forming fungi have succeeded in correcting this fundamental omission in an indirect manner. 4. There is no evidence in favour of the supposition that the Myxogastres are degenerated members of the vegetable kingdom, whereas the idea that fungi originated by differentiation from chlorophyll-bearing plant ancestors is generally admitted. 5. The Myxogastres with allied forms included by Zopf differ from all known members of the vegetable kingdom in having the component cells naked, or without a cell-wall, during the entire vegetative phase. Classification. In addition to the Myxogastres as defined in the present work, other smaller groups, as the Acrasicae of Van Tieghem ; the Monadlncae of Cienkowski, &c., are supposed to possess certain important characters in common with the Myxogastres, and have been by some authors united into one group. The following outlines of the various schemes of classification will indicate the views of affinity as proposed from time to time. The term Mycetozoa as already explained, was first used by De Bary, 1 and in his latest work on the subject this name is retained as follows : — 1 D'tc Mijcetozoeii, ZclischriJ'i f. nnss. Zoohxjic, Bd. .\. 18.09. 22 A Mo?ioff)-ap/i of tl/e Myxogastres. MYCETOZOA. I I . I Swarm-cells coalescing to Swarm-cells becomiiif; form a plasmodium. aggregated, but not = Myxomycetes. coalescing to form a Plasmodium. = ACRASIEAE. Cienkowski's Nnclcariae and Vampyrellae along with such genera as Bur sella, Protomyxa, Myxastrum, Monas, Monadopsis, Pscudospora, Colpodella, and Plasmodiophora are considered by De Bary as " doubtful Mycetozoa " for the following reasons. " I here exclude from the ranks of the true Mycetozoa a few forms or groups of forms, some of which have been occasioually mentioned in the preceding sections. Those forms, so far as they are known, have many points of resemblance with the Mycetozoa, but either our knowledge of them is imperfect, or else they depart so far in certain points from the typical Myxomycetes and Acrasieae, that it is better to leave their position in the system for the present undetermined." ^ Although De Bary clearly indicated that the existing classifi- cation of the Mycetozoa was very imperfect, being the outcome of pocket-lens observations oa mature forms, and as above indicated, pointed out a scheme in accordance with the modern system of research, yet, lacking time, he never completed all the details necessary for a thorough revision of the group. This was done however by Dr. Rostafinski, a student of De Bary's, and as at least some of Rostafinski's important work on this subject was done by Rostafinski in De Bary's laboratory at Strassburg, we may presume that the general scheme was more or less inspired by De Bary. Rostafinski confined his attention to the group Myxomycetes as defined by De Bary, and this group he calls Mycetozoa, thus using the same name in a narrower sense than De Bary, who included the Acrasieae that do not form a plasmodium, and in his ilonograph ^ recognized two primary divisions — ExosPOREAE, 1 Fiouji Mycetozoa ciud Bacteria, Engl. ed.. p. 446. - Monyrajia Sluzowce, Paris, 1875 (in Polisli). .1 Monograph of the Myxogastres. 23 having the spores produced externally on slender sporophores, a feature met with only in the genus Ccralmm ; and Endosporeae, having the spores produced in sporangia. In the second " Aj^pendix " to the Monograph, amongst other changes in the classification, the Exosporeae division is omitted altogether, and the Eudosporous division alone retained. In Rostafinski's classification the two primary divisions are founded on the colour of the spores as seen under the microscope by transmitted light; each primary division is further divided into two sections determined by the presence or absence of a capillitium as shown in the following arrangement — MYCETOZOA. ENDOSPOREAE. Amaurosporeae. Lamprosporeae. Spores violet or brownish- Spores yellow or brownish, violet. never violet. , \ I Atrichae. Trichophorae. Atrichae. Trichophorae. Von Tieghen uses the name Myxomycdes in a broader sense than De Bary's Mi/ccf ozoa, ciud arranges the groups as follows: — ^ MYXOMYCETES. THALLUS. I Unicellular. Pluricellular. without a Plasmodium. _ I = Plasmodiophoreae. r 1 I I cells aggregated but not fusii to form a plasmcdinni. I =Acra.sieae. Cells fusing to form a Plasmodium. ^ ! ^ Spores produced in Spores produced externally, a sporangium. =Ceratikae. = Endoiiyxeak. 1 Traitcdc lJu(on>\ji',; p. 992 (1884). 24 A Monogra2)h of the Myxogastres. Zopf s Mycdozoa include many forms that do not come within De Bary's conception of the Mycdozoa as already defined. The following illustrates the primary divisions as adopted in his latest work.i A. MONADINEAE. Mostly aquatic, partly parasites ; resting cysts are usually formed. I. Monaclincae azoosjjorcae. Vampyrelkac ; Bursellineae ; Monocystaccae. II. Monad-ineae zoos2}oreae. Pscudos]Joreae ; Gymnococcaccae ; Flasriiodio2Jhoreae. B. EUMYCETOZOA. Aerial. Never parasites ; plasmodia always present, generally well developed ; fructification generally well developed. I. SOROPHOREAE. Gfcttulincae ; Didyostdiaceae. II. Endosporeae. a. PcritricJieae. Clcdhroptydiiaccac ; Crihrariaccae. /3. Endotricheae. * Stereonemeae. Cahariaccae ; Amaurodiadaceae. * * Ccdonemcae. Trichiaccae ; Arcyriaccac ; Bdicidariaceae ; Liceaceac ; Fcri- chaeTiaccae. III. EXOSPOREAE. In Raunkier's Myxomycdcs Daniae - the following classification is proposed — 1 Die Pilztliiere ocler Sclileiinpilze (Sclienk's Handbnch der Butanik, ill Encylcl. der Naturtoisscnuchaften), 1884. 2 Botanisk Tiddskript, 17 Bind. (1888). A Monograph of the Myxogastrex. 25 A, Capillitium absent. I. HOMODERMEAE. Liceaceae. {Ttthulina, LindUadia.) II. Heterodermeae. Clathroptycliaccae. {Enteridium, Clathroptychium.) Cribrariaceae. {Crihraria, Dictydium.) B. Capillitium present. III. C(ELONEMEAE. Arcyriaceae. {Pcrichaena, Zachnoholus, Arcyria, Cor- nuvia, Lycogala) Trichiaceae. {Hcmiarcyria, TricJda.) IV. Stereonemeae. Physaraceae. {Badhamia, Physarum, Tilmadochc, Fuligo, Leocarpus, Craterimn.) Didymiaceae. {Chondriodcrma, Lcpidoderma, Didymium, Spumaria.) Stemonitaceae. {Lamprodcrma, Bncrthcncma, Ancyro- phorus, Comatricha, Stemonitis, Brcfeldia, Ecticularia.) The Myxomycetcs are divided by Schruter^ into three prin- cipal groups as follows — A. Mature fructification consisting of a mass of free spores. * Saprophytes ; the amoeboid bodies unite in masses, but do not coalesce = Acrasieae. * * Parasitic in the interior of living cells, forming in the known instances, a true plasmodium = Phyto- myxineae. B. Spores formed in the interior of sporangia, or on the outside of discoid or columnar fructifications = Myxo- GASTRES. At the close of the vegetable period, the passage of the motile Plasmodium into the stationary reproductive condition is abrupt, and takes place as follows; the surface of the plasmodium becomes elevated into one or usually many protuberances, the original investment of the plasmodium is continuous over these 1 Eiigler u. Prantl.s' Natiirl. Pflaiizenfam., 30 Leif., von Y. Sclirolor, 1889. 26 A Monograph of the Myxogastres. protuberances into which the whole of the protoplasm passes, leaving behind the remainder of its pellicle attached to the substratum, and known as the hypotlialhcs. When these protuberances, which may be sessile or stipitate, are symmetrical and individually distinct, they are called sporangia; when sporangia are irregular in form, usually veinlike and creeping, the term jjifi-^inodiocarp is used ; finally, when the sporangia are densely aggregated, so that their individuality disappears to a greater or less extent, an acthlium is produced. The three conditions are connected by intermediate links. Aethalia are most fi'equently sessile on a broad base, as in Tubulina cylindrica and Entcridium olivaceiim, but stipitate aethalia are not un- common, especially in the Trichiaceae, where the transition from tj-pical sporangia to aethalioid forms in many species is very instructive. As previously stated, the late Professor De Bary was the first to show, by his admirable researches on the morphology and physiology of the Myxogastres, that the systematic arrange- ment then in vogue was no longer tenable, owing to the fact that it was founded on analogies rather than affinities; and although at the present day it must be admitted that, within the group, affinities are far from being settled, yet, the appear- ance of Dr. Rostafinski's Monograph based on De Bary's I'e- searches, gave a fresh impetus to the study, and showed in a masterly manner, that well-marked morphological features, far beyond the ken of pocket-lens revelation, could be utilized in connection with the systematic disposition of the members of the group. Among the essentials still lacking for an approxi- mately correct classification are more especially — amount of variability, as also its direction in the various sections ; the relative value of such structures as the columella, capillitium, sporangial, aethalioid and plasmodiocarp forn;s; the presence or absence of lime, as also its amorphous and crystalline condition as presented in different sections ; and finally, what is undoubt- edly of primary importance, a complete knowledge of the life- history of at least the representative species of each section ; and although complete life-histories might not in all cases be A Mo?io(jrrfp// of tlie Myxogastres. 27 possible, yet iu so difficult a subject, the modus o'j)crandi alone would enable others to follow along the same lines. In the present state of knowledge, the markings usually present on the epispore appear to be constant within narrow limits, are often very characteristic, and of value as one factor in the discrimina- tion of what we at present consider to be species, nevertheless I wish to express the strongest dissent to species founded on spore characters alone, or indeed on any one character. Rostafinski was the first to use spore characters in a specific sense. It is important to bear in mind, that the apparent nature of spore ornamentation depends entirely on the amount of magnifying power used; the complex epispore of Trichia affinis appears only as a confused series of minute irregular projections under a quarter-inch objective, consequently there is no absolute char- acter in the spore unless a uniform magnifying power is agreed upon. The objective used in determining the nature of the epispore as described in the present work is a ^^ oil immersion, which gives a magnifying power of 1200 diameters. So far as the synonymy is concerned, I can only repeat in substance what I have written on a previous occasion in connection with the same subject. I have not included the synonymy further back than Rostafinski's Monograph, unless justified by the presence of type or authentic specimens. Rostafinski has given syno- nyms dating from the time of Micheli (1729), but on referring to the earlier descriptions, I realize my weakness, in not being able to reconcile the wonderfully brief descriptions, and equally crude drawings, with modern species, which in many instances reipiire a magnifying power of 1000 diameters for their deter- mination, hence I wish to be clearly understood that the synonyms headed "Rostafinski's Synonyms," are copied from Rostafinski's Monograph without any attempt at corroboration. I feel certain that nearly one-third of Rostafinski's work would not have been sacrificed to synonyms unless they mean some- thing more than I have been able to discover, hence I have not felt justified in ignoring them altogether. 28 A Monograph of the Myocogastres. MYXOGASTRES (Feies). Myxogastres, Fries, Syst. Myc, iii. p. 67 (1829) ; Schroeter, Kr. Fl. SMcs., Vol. iii. p. 98. Myxomycetes, Wallr., Fl. Cryi^t., ii. p. 333 (1833) ; Cooke, Brit. Myx., p. 1 ; Raunk, Myx. Dcm., p. 20 ; Sacc, Syll., Vol. vii. pt. i. p. 323. Mycetozoa, De Bary {Die Mycctozocn, 1864); Kost. {Mon. Sluzoivce) (in part). Pilzthiere, Zopf, in Schenk's Handb. der Bot., Vol. iii. (1884) (in part). Consisting during the entire vegetative period of an accumula- tion of nahed cells forming a plasmodiuiii possessed of the poiver of movement, and invested with an imperfectly differentiated, yielding mcmhrane. This plasmodinm at length hccojiics trans- formed into sporangia containing numerous spores lohich form a powdery mass at maturity. First product of spores on germination either ciliated zoospores or amoeboid cells, which after repeated bijKirtition and conjugation, combine to form a p)lasmodium. The species are usually aerial, the plasmodium stage being passed in rotten wood or amongst decaying vegetable matter, and coming to the surface only to form the sporangia ; never- theless several species have been observed to pass the whole of tlieir existence under water. The sequence from the aquatic to the aerial condition may frequently be seen in the same batch when developing on logs that are partly submerged. The most pronounced feature in the evolution of the Myxo- gastres is in connection with spore dissemination, and the following arrangement is based on the relative development of the capillitium, which is seen in its most perfect form in the genera Trichia and Arcyria. In the Gastromycetcs I have shown else- where ^ that the sequence of evolution all tends towards securing in the most efficient manner, the dissemination of the spores ; commencing with the subterranean genera Bhizopioggn and ' A Monorimph of the British Gastromycetes. Ann Bot, Vol. iv. pp. 1—103 (4 plates). A Monograph of the Mii.rogastreif. 29 ITymcnogastcr, where the spores are liberated only by tlie decay of the plant without any apjoarent morphological provision for their dispersion, we trace, as the above ground genera appear, the gradual evolution of the capillitium through such genera as Diploderma and Scleroderma, until in Lycopcrdon we find the highly specialized hyphae forming the capillitium effecting at maturity the rupture of the peridium, and consequent disper- sion of the spores. This phase is the most perfect, so long as dispersion is effected by mechanical methods, but in the Phal- loidcae we find the mechanical method superseded by a totally different idea ; brilliant colours, usually combined with a strong odour, and the presence of a sweet substance, bid for the visits of insects, which in return disperse the spores. In the Myxo- gastres the mechanical method of spore dispersion is the most perfect condition attained, as illustrated by the rough, elastic capillitium of many species of Arcyria. The above allusion must not be interpreted as implying relationship between the Gastromycetes and the Myxogastres ; in fact I see in this point of agreement no more proof of affinity than exists between the Pballoideae and those flowering plants that make use of the same announcements to insects, in the form of colour, scent, and honey in connection with pollination. I consider the Myxogas- tres as illustrating one of the earliest known attempts at differentiation in the direction that has eventually resulted in the mass of organisms constituting the vegetable kingdom, but having originated from the Flagellatae, a group more in touch with the animal side of life, the work of developing individuality has been slow, as illustrated by the tardy appearance of cellu- lose cell-walls, which, as would be expected, is most complete in the newly evolved reproductive phase, itself to a great extent the outcome of a gradual change of environment from aquatic to aerial — but the radical mistake, after having adopted the plant line of development, consisted in the non-development of chro- matophores, and retention of the animal mode of nutrition, which in the plant world means parasite or saprophyte. The fungi, a later group, differentiated from ancestors that had already evolved the leading plant characteristics, including cell- 30 A Monograi^li of the Miixogastres. walls, chlorophyll, starch, hence in this respect are more typical plants than the Myxogastres ; but in the fungi the check to pro- gress was due to the degeneration of chromatophores, already evolved by their ancestors, whereas in the IVIyxogastres the check was due to their inability to differentiate these essentials. MYXOGASTRES. Order. Peritricheae. Wall of sporangium not incrusted with lime; capillitium absent or formed from wall of sporangium. Sub-Order. Tiilmlinae. Wall of sporangium not perforated. Sub-Order. Cribrariae. Wall of sporangium perforated. Order. COLUMELLIFERAE. Wall of sporangium without lime ; capillitium originating from a central, usually elongated columella. Sub-Order. Stcmonitac. Capillitium springing from every part of an elongated columella. Sub-Order. Lctm'prodcrmae. Capillitium springing from the apical portion of a short or elongated columella. Order. LiTHODErxMEAE. Wall of sporangium with an external deposit of lime; capillitium present. Sub-Order. Didymcae. Threads of capillitium without lime. Sub-Order. Physarac. Capillitium containing lime. Order. Calotricheae. Wall of sporangium without an external deposit of lime ; capillitium present, not spriDging from a columella. Sub-Order. Trichcac. Threads of capillitium free, simple or branched, not anastomosing to form a network. Sub-Order. Arc/jriae. Threads of capillitium either attached by one end with the free tips more or less branched, or combined to form aji irregular network. The following diagrammatic arrangement is an attempt to illustrate my ideas of relationship between the several orders. The bracketed generic names represent the 2:)oints of contact A Monograj^h of the Myxogastres. :u between respective orders, and the names in italics to the riglit and left indicate tliose genera that illustrate the maximum of development characteristic of the orders to which they are respectively attached. Choiidriudenna. Badhamia. j_ -LiTHODERMEAE. 1 '' / Diac V Laiiipr laca. \ xlurma./ Stemonitis. Eoierthonema. 1 COLUMELLIFERAE. . 1 t /Siphot (, TnL ychiuniA ulina. / Crlhrai ia. Dicfydixm. 1 PF.RTTRTnHKAR i ((^latliroptj^cliimiiA Pericliaena. / Trie! hi a. .Calotkicheae . Arcyria. i In each order we find the special characteristic idea evolving through a sequence of genera, the terminal one not connecting with any higher order, hence the special feature terminates abruptly within the order where it originated, and it is invari- ably in some comparatively undifferentiated genus near the initial point of each order that we meet with the suggestion of a new line of evolution, which at its maximum of development, constitutes the characteristic feature of the order immediately 32 A Monogra2)h of the Myxogastres. in advance of the one from which it emanated in an incipient condition; hence in the Feritricheae, starting with the genus Tuhulina, the marked characteristic of the order — a perforated sporangial wall — evolves gradually through Frotodermmm to Didydimn, where it attains its highest phase of development, and then terminates abruptly ; whereas' in Entcridmm, a com- paratively primitive genus, we get a transition from slightly perforated walls to a complete breaking up of their substance into an irregular network, which leads by degrees to the central columella and capillitium characteristic of the Colvyinellifcrae. FERITRICHEAE. The present order includes the most primitive types met with in the Myxogastres, illustrated by such genera as Ttilulina, of which the plasmodiocarp condition, representing a plasmodium covered by a common cortex, is the lowest phase ; when the substance of the plasmodiocarp becomes broken up into smaller portions by special walls, an aethalium is produced, and finally, by the gradual isolation of the components of an aethalium we get the sporangium or most highly specialized form of repro- ductive organ, characteristic of the higher sections ; but we everywhere meet with plasmodiocarp and aethalioid conditions, proving the comparative instability of the acquired sporangial phase. In the present order we learn that the primitive idea in connection with spore dissemination was that of modifying, or rather differentiating, the sporangial wall. The original function of the wall was protective, as illustrated in the genus Tuhdina, where we find this to be its only function, there being no provision for spore diffusion other than that brought about by the decay of the wall, which does not take place readily, and the subsequent dispersion of the spores by wind or rain. Passing on to such genera as E^iteridium and Clalhro^itychium, we find the sporangial wall yet perfonning, as it continues to do in every section, the function of protection, and remaining intact until the spores are fully differentiated ; but in these genera certain portions of the wall have become specially A Afonorjnrph of the Myjroga^tres. 33 thickened, and at the period of maturity, the thickened portions are persistent, while the remaining portions of the wall dis- appear, and as this thickening is usually of a determinate nature, and utilized as a generic factor, we meet with the small, fairly regular openings in the lateral walls presented by one of the members of the genus Entcridium, or the still larger openings of Clathroptychium ; but, although the modification is a step in the right direction, owing to the fact that the species constituting the two genera named retain the aethalioid con- dition, the new idea is of but little practical value; and it is only when we come to Crihortria, the components of Avliich for the most part have reached the condition of forming small detached Plasmodia, each only sufficiently large to form a single spor- angium, that we realize the value of the perforated sporangial wall, which, other things being equal, is better than the sealed up, entire sporangium, inasmuch as at the moment the spores are mature, there is a chance of being removed from the sporangium by wind or rain ; but in Crihraria the stem, although usually elongated, is rigid, and it is in the genus Didydium that arrangements for spore diffusion from a perforated sporangium is most perfect. In this genus the thickened, permanent portions of the sporangium are arranged in the form of ribs radiating from the base and converging again at the apex, being connected by very thin, transverse bars, forming a hollow sphere of lattice-work ; this globe is pendulous from a long stem curved at the apex, and while the stem is rigid during the young 'condition, towards maturity the upper portion above the curve becomes shrivelled and flaccid, and the sporangium hangs in a condition ready to be dangled by the wind or the movements of minute insects ; how effectual this method of spore diffusion is in its way is proved by the specimens being generally found destitute of spores. In Entcridium, as already mentioned, we find in one species the wall symmetrically perforated, whereas another species has the walls of the component sporangia forming the aethalium so irregularly perforated, and forming a variable network, that at first sight the aethalioid nature of the mass is masked, and this species may be considered as forming D 34 A Monograph of the Myxogastres. a connecting link with another order, the ColumcUifcme, where the caiiillitiura originates in a totally different manner. The following arrangement illustrates the evolution of the genera included in the Pcritrichcac, and the points of affinity with other orders — DiCTYDIUM. I Cribraria. Clathroptychium.- ft o -Prutodermium. PERITEICHEAE. (British genera are printed in small capitals, exotic genera in italics.) ANALYSIS OF THE GENERA. PERITEICHEAE. Sub-sect. TUBULINAE. TUBULINA. Sporangia crowded to form an aethaUum. , Protodermium. Sporangia scattered. Sub-sect. Cribrariae. Orcaddla. Sporangia perforated at tlie apex. Enteridium. Sporangia with small lateral perforations, or broken up into an irregular network. Clatiiroptychium, Sporangia with large symmetrical open- ings in side walls, apices of sporangia permanent. Cribraria. Permanent portions of upper part, or whole of sporangium forming an irregular network. DiCTYDIUM. Permanent ribs of sporangium radiating from the calyculus and converging towards the apex, joined by thinner transverse bars. Tuhalina. 35 Sub-sect. TUBULINAE. TuBULiNA, Pers. (emended). Sporangia scattered and regular, or irregular and forming a plasmodiocarp, or compacted together to form a naked or corti- cated aethalium ; walls very thin, single, often iridescent, not perforated ; columella and capillitium absent. Tuhuliim, Persoon, Syn. Fung., p. 197; Rost., Mon., p. 219; Cooke, Myx. Brit., p. 54 ; Sacc, Syll., Vol. vii. p. 406 ; Schroeter, p. 102; Zopf, p. 172 f. Licca, Schrader, Nov. PI. gen., p. 17 ; Rost., Mon., p. 218; Schroeter, p. 102 ; Zopf, p. 171 ; Sacc, Syll., Vol. vii. p. 404 ; Cooke, Brit. Myx., p. 54. Lindhladia, Fries, Summ. Veg. Scand., p. 449; Rost., Mon., p. 223 ; Cooke, Myx. Brit., p. 55 ; Sacc, Syll, Vol. vii. p. 408 ; Zopf, 172. A careful examination of ample material has convinced me that the three genera given above constitute but one natural genus, and Tuhidina, having priority, has been adopted. The structure and origin of an aethalium being unknown to the old mycologists, explains the origin of these genera ; but it is remarkable that Rostafinski, who so lucidly explains the origin and value of aethalia, did not notice the sequence so clearly stated in his own diagnoses of the three genera, which he retains. In all three the sporangial walls agree in being membranaceous and not perforated ; Licca has the sporangia regular, as in L. minima, Fr,, or irregular and forming a plasmodiocarp, as in L. flexuosa, Pers. ; in Tulmlina the sporangia are cylindrical or prismatic from mutual pressure, and combined to form a naked aethalium ; while in Lindhladia, the sporangia also form a naked aethalium, but although prismatic, as in Tuhulina, arc not quite so much elongated ; finally, the species described as Licca simmarioidcs, Cke. and Mass., belongs to one of the trio, but differs from all in having the sporangia flexuous, and in the aethalium being covered with a common cortex. If Rostafinski's view as to the autonomy of the three genera in question is correct, it follows that the aethalioid condition of such species as Hcmiarcyria ruhi/ormis, Rost., must not only be separated from the form with simple, normal sporangia, 36 A Monograph of the Myxogastves. but must constitute a new genus, which exactly illustrates the difference between Licca and Lindhladia or Tuhdina respectively ; whereas the only distinction given by Rostafinski between T^ihclina and Lindhhulia amounts to saying that in the former the sporangia are elongated ; in the latter, shorter. A well-defined hypothallus is often present, and in some species, as T. stipitata, B. and C, forms an elongated, stem-like stmcture which possibly consists of sterile, modified sporangia. Dr. Geo. A. Rex has kindly furnished me with a sequence of specimens, all belonging to Tululina cylindrica as defined in the present work, which to my mind show clearly the transition from Tiibulina to Siplioiotychium, and thus furnishing a connect- ing link between the Fcritrichcae and the Culumelliferae. Distrih. Europe ; North America ; India; Ceylon; Australia; Bonin Island: species, 11. A. Sporangia, regular or plasinodiocarp. Tubulina minima, Mass. Sporangia scattered, sessile on a broad base, hemispherical or varioicsly elongated, little depressed ; wall thin, smooth ; from clear to ohscure hrown; torn above into ^e,\evsl persistent, irregular teeth ; mass of spores umber, spores globose, minutely warted, 10 — 12 /x diameter. Licca minima, Fr., Syst. Myc, iii. p. 199 ; Karst., Myc. Fenn., iv. p. 125; Sacc, Syll., 1385. Licca variabilis, Schrad., Nov. pi. gen., p. 15, t. vi., £f. 5, 6. Uxsicc. — Karst., Fung. Fenn., 798. (Specimen from Fries in Herb. Berk.) Britain (Bulmer, Yorks). On pine wood and sawdust. Germany ; Sweden ; Finland. An obscure species, which, from its scattered habit, is easily overlooked ; when circular from "5 — 1 mm. across ; frequently 2 — 8 mm. when elongated. The colour varies from almost chest- nut to blackish brown ; ruptured above in an irregularly stellate manner. The inner surface of wall of sporangium has numerous coloured cells or masses arranged in groups. Tubulina. 37 Rostafinski's gratuitous suggestion that the present species is a Phclonitis is altogether wrong. Tubulina flexuosa, Mass. Sporangia scattered, sessile on a broad base, circular or varioushf elongated and flexicous, wall thin, single, shining; deep olive or tinged loith hroiun ; mass of spores olive ; spores globose, pale olive-brown, minutely wartcd, the warts showing a tendency to form very short ridges, 12 — 15 [x diameter. Licea fleoMOsa, Pers,, Syn. fung., p. 197, t. 1, f. 5, 6; Rost., Mom, p. 218; Schroet., p. 102; Sacc, 1382. (Specimen named by Rostafinski in Herb. Berk.) On decaying wood, especially pine. Britain (Aboyne, N. B.) ; Finland ; Sweden ; Germany. From 1 — 1"5 mm. when circular, often 2 — 3 mm. when elongated. The wall of the sporangium is furnished inside with a layer of fine granules of organic matter. (Rostafinski's Synonyms.) Licea fiex^iosa, Pars., Syn. p. 197, t. 1, f. 5, 6 (1801). Tululina flexuosa, Poir., Enc, I.e. viii. no. 8 (1808). L. serpula, Fr., Sym. Gast., p. 12 (1817). L. spadicea, Fr., cfr., I.e. iii. p. li)7 (1829). Lycogala contortium, Lk., Herb, non Ditm. Acthalium minimum, Wallr., in litt. Aethalmm microscojncum, Wallr., Herb. Zicea ahUacca, Wallr., Fl. Ger. cr. n. 2106 (1833). Tubulina spermoides, Mass. Sporangia cylindrical, shorter than broad, crowded, rarely scattered towards the circumference of the clusters, apex slightly rounded, wall very thin, dull, breaking up into whitish fragments, seated on a well-developed, thin hypothallus; mass of spores olive ; spores globose, 6, 7 /x diameter, equally covered, cdl over with minute iveirts. Licea spermoides, B. and C, Grev. v. 2, p. 68. Crihraria argillacea, Sacc, Syll., 7, n. 1415 (in part). (Type in Herb. Berk. n. 10854.) On wood, leaves, &c. Alabama. Sporangia in a single stratum, 38 A Monogr<(2>li of the Myjcogastyes. often forming patches 6 — 9 cm. long and broad ; the hypothalkis although thin, is tough, and can be peeled otf the matrix. Sporangia about 5 mm. diameter and not quite so high ; before dehiscence yellowish olive and resembling a crowded Ferichaena. Tubulina speciosa, Speg. Sporangia densely gregarious, fasiculate, fascicles subcircular, 1 — 2 cm. long and broad, 1 — 1 mm. high, seated on a slightly convex hypothallus ; sporangia cylindrical, apices rounded, dehiscing irregularly, at fio^st ficsh-colonr then cinncibar, shining ; spores warted, rusty-ochre ; depressed, dry concavo-convex, with a thick ish margin. Spegazzini, Nov. Add. ad Myc.Ven. n.l23; Sacc.,Syll.,7,n.l393. On rotten wood of Finns silvcstris. N. Italy. I have had no opportunity of examining the present species, the spores of which are described as concavo-convex when dry, but in all probability become spherical when moist. Tubulina stipitata, Rost. Hypothallus stem-like, often sulcate, spongy within; spo- rangia crowded in a sub-globose head, cylindrical, apices rounded ; walls very thin, evanescent ; mass of spores brownish- umber ; spores globose, 4 — 5 'ix diameter, about three-fourths of the surface luitJi a regular small nctwm^k, the remainder with much larger mesltes. Rost., Mon., p. 223, f. 2; Cooke, Myx. Brit., f. 2; Sacc, Syll., 7, n. 1392. Lieea stijntata, B. and C, Grev. v. 2, p. 68. (Type in Herb. Berk. n. 10851.) Exsicc. Fung. Cub. Wrightiani, 551. On dead wood, bark, &c. Stem 2 — 4 mm. thick and high, umber, porous within; sporangia 1"5 — 2 mm. long. The sjiores are globose when thoroughly soaked. Dr. G. A. Rex of Philadelphia has recently sent specimens of the present species having the hypothallus very much reduced, and also informs me that sessile forms are not un- common, hence the specific distinction between 2\ cylinchica and the present species turns mostly on spore differences. Cuba; Java; Bonin Islands; United States. Tahulina. 39 Tubulina guaranitica, Roumeg. Gregarious ; sporancjia glohose, sometimes slightly attenuated below on passing into the stem; wall single, thin; clingy ochraceous with a tinge of olive, sometimes greyish below, about 1 mm. diameter; ston rather thick, slightly incrassatcd doivmoards, and 2^a!ising into a hypothalhis, 1-5—2 mm. long, ochraceous-brown ; mass of spores umber ; spores globose, smooth, 7 — 8 [X diameter. Tiibulina guaranitica, Roumegu^re, Fung. Sel. exsicc., no. 5196. Licea ? guaranitica, Speg., Fung, guaran.. Pug. I, no. 322. On decayed bark. Guarapi ; Argentine Republic. With the general appearance of a small specimen of Ttdmlina stipitata, differing in having only a single sporangium, or aethalium surrounded by a thin, friable cortex. The dimorphic capillitium described by Spegazzini as consisting of septate threads is clearly nothing more than the mycelium of some minute fungus spreading in the mass of spores. The tail-like appendages to the spores mentioned by the author are to be explained in the same way. The above description is drawn up from the specimens in Roumeguere's exsicc. These speci- mens were supplied by Balansa, and may therefore be con- sidered as authentic, inasmuch as Spegazzini's Fungi guaranitica is an account of the Fungi collected by Balansa. B. Acthaliuvi nalrd. Tubulina cylindrica, Rost. (f 1 — 0). Sporangia usually crowded and forming a naked cake-like brown aethalium, rarely scattered, seated on a well-developed flat or slightly convex hypothallus, cylindrical or subangular from mutual pressure, more or less elongated, apex rounded ; wall single, thin, often iridescent; mass of spores brownish- umber; spores globose, ahoat three-fourths of the surface with a regular, Jine-meshcd nctworh, the remainder with sccdtered ridges and ivarts, 7 — 9 \x diameter. Rost., Mon., p. 220 ; Ckc., Myx. Brit., p. 54; Sacc, Syll., 1391; Schrocter, p. 102. 40 A Monograph of tiff Mi/rogastres. Tuhdina nitidissima, Berk., Journ. Linn. Soc, v. 18, p. 387 (from examination of type); Licea rulifm-mis, B. and Curt, (from exam, of type). Licca micro&perma, B. and C. (from exam, of type). Exsicc. — Syd., Myc. March., 1498; Cke., Brit. Fung., ed. ii., 528 ; Karst., Myc. Fenn., 697 ; Fekl., F. Rhen., 1470 ; Roum., Fung. Gall, 3066 ; Ellis and Everh., N. A. Fung., ser. II. 2096. On wood. Forming compact cakes varying from half an inch to a foot or more in diameter, varying from umber-brown to chestnut, the surface granular with the convex apices of the sporangia. When immature and pulpy, of a beautiful straw- berry colour. Hypothallus spongy, resembling when thick the sterile base of a Lycopcrdon. Dr. Rex considers that two distinct species are confounded under the above name. Britain (Apethorpe, Powerscourt, Brandon, Scarboro', Car- lisle, Glamis, N. B.) ; Germany ; France ; Sweden ; Finland ; Hungary; Belgium; United States; India; Ceylon; Australia. (Rostafinski's Synonyms.) Tululifcra cercdum, Fl. Dan., t. 659, f. 2 (1777). Tubidifcra arachnoidea, Jacq., Misc., t. 15 (1778). Mucor iuhulosus, Retz. Stemonitis ferruginea, Batsch, f. 175 (1786). Sphacrocarpus cylindrims, Bull., t. 470, f. 3 (1791). Sphacrocarpnis fragiformis, Bull., t. 384 ? (1791). Tuhiferaferruginosa, Gmel., Syst. 1472 (1791). Tiibifera cylindrica, Gmel, Syst. 1472 (1791). TuUf era fragiformis, Gmel, Syst. 1472 (1791). Trichia fragifm-mis, With., Arr., iii. 480 (1792). T'lbhidifcra coccinea, Trent., p. 243 (1797). Licca tuhulina, Schrad., N. G., 16 (1797). Licca clavata, Schrad., N. G., 18 (1797). Tuhdina fragiformis, Pers., Disp., p. 11 (1797); Pers., Syn., t. 4, f. 3, 4. Tnlndimtfallax, Pers., Obs., ii. 28 (1799). Tuhulina fragiformis, a. papiUata, /3. clavata, y. coiiica, b. coccinea, e. opcrculcda: Pers., Obs., ii. 29. Tuhulina. 41 lieiicularia multiccqmda, Sow,, t. 179 (1790). Tuhidina cylindrica, D, C, FJ. F,, 674 (1805). Tuhdina fragifera, Poir., Ency., viii. No. 3 (1808). Zicea fragiformis,'Nees,f. 102 (1816); Eng. Fl., v. p. 821; Cooke, Hdbk., 1194; Fung. Brit., ii. 528. Bermodium fallax, Nees, f. 103 (1816). Licca cylindrica, Fr., S. Myc, iii. 195 (1829) ; Eng. FL, v. 391 ; Cooke, Hdbk., No. 1193. Licea iricolor, Zoll., in Flora (1847), p. 300. Tuhulina concjlohata, Preuss., Linnaea, 140 (1851). Tuhulina effusa, Mass. ActJiali'itm naked, sporangia seated on a firm, common hypo- tliallus, irregular from mutual pressure, in a single stratum or superposed, walls very thin, lustrous, grown together ; free apices of spoQ^angia slightly convex, giving the surface of the aethalium a granular appearance ; mass of spores ochraceous-umher ; spores globose, yellowish-brown, very indistinctly vcrruculose, 6 — 8 ix diameter. Lindhladia effusa, Rost., Men., p. 223; Schroct., p. 103; Cooke, Myx. Brit, p. 55; Sacc, Syll., no. 1395. On the ground and on wood. Britain (Leicester, Epping, Scarboro', Aboyne, Forres, Linlithgow); Germany; Sweden; France ; Bohemia. Often forming compact, flattened cakes extending for three or four inches, sometimes much smaller; superficially closely resembling Tuhulina cylindrica, but the mass of spores with a yellower tinge, and the present species also differs in the shorter sporangia, which are not so uniform in their arrange- ment as in T. cylindrica. (Rostafinski's Synonyms.) Licca effusa, Ehr., Sylv., p. 26, f. i. (1818). Aethalium melaenum, Chev. Byss., iii. (1837). Lindhladia tuhulina, Fr., S. V. S., 449 (1849). Acihiliiiiii alrum, Preuss., Linnaea, 141 (1851). 42 A Monoijra])li of the Myxogastres. Reticularia maxima, Corda, Ic. vi. 85 (1852), Beticularict granulosa, Qi^rstd. MSS. 0. Aetlialium covered with a common cortex. Tubulina spumarioidea, Cke. and Mass. Aethalium large, irregular, cortex membranaceous, higUt grey, with anastomosing veins inside ; walls of sporangia almost colourless, very thin and soon disappearing; mass of spores orange with a tinge of olive ; spores globose, with oxtther large, oltuse loarts, 16 — 18 /x diameter. Licea sj)%imarioidca, Cke. and Mass., Grev., vol. xvi. p. 74 (1888). Running over twigs and on the ground. Near Melbourne, Australia. Aethalia subglobose or irregular, varying from | to 1 inch across, creeping up grass and twigs like Siyivmaria alhcc ; common cortex bright grey, thin, subpersistent, with numerous anasto- mosing vein-like thickenings in the inside of the wall. Spor- angia densely crowded, flexuous, wall thin, at first imperforate, soon disappearing. Imperfectly known species. Tubulina brunnea (Preuss.). Closely gregarious; sporangia ochraceous-brown, rotund, sub- depressed, wall papyraceous, irregularly torn, upper portion disappearing in a subcircumscissile manner ; spores globose, minute, ochraceous, threads none. Licea hrunnca, Preuss., Hoyersw., n. 275 ; Schroet., p. 102 ; Sacc, SylL, 1887. On semi-putrid trunks of pines. Germany ; Silesia. Tubulina Lindheimeri, Mass. Broadly effused, externally blackish, mass of spores ferru- ginous-orange ; spores globose, smooth, — 7 fx diameter. Luxa Lindheimeri, Berk., Grev. v. 2, p. 05 ; Sacc, Syll., v. 7, n. 1388. Tuhulina. Pvotodenmum. 43 (Type in Herb. Berk. n. 10848.) On dead bark. Texas. The type specimen is so thoroughly crushed that it is im- possible to say whether the plant is a Licca or a Tuhulina, but in all probabiKty the latter, as there appear to be indications of the bases of crowded sporangia. Tubulina caespitosa, Mass. Peridia aggregated in tufts or clusters, crowded, sessile, smooth, brown or blackish-brown ; spores dingy ochre, smooth, globose, .00025 in. in diameter (=5.5 ii). Thysarunt cacsintos%tm, Peck, 26th Report, State Mus. N. York, p. 75. Rotten wood. Greenbush ; U. States. Protodermium, Rost. Sporangia simple, solitary, regular in form; mass of spores blackish violet ; capillitium entirely absent. Protodcrma, Rost., Mon., p. 00; Schroeter, p. 103; Cooke, Myx. Brit., p. 10. Frotodcrmium, Sacc, Syll, Vol. vii., pt. I., p. 325. Rostafinski's name ProtGderma was altered by Berlese in Saccardo's Sylloge to Protodermium, as the former had been already used for a genus of algae. The present genus is identical with Talmlma except in the colour of the spores, a character which I look upon with doubt. Colour as presented by spores is undoubtedly of some signifi- cance, but being altogether in ignorance as to its relative physiological value, I cannot accept colour in Rostafinski's sense as being of such supreme importance as to constitute the basis for the primary divisions of the Myxogastres, even if the divisions indicated had really existed in nature, which however is far from being the case. Disirih. Europe; United States: species 1. Protodermium pusillum, Rost. Sporangia gregarious, distinct, hemispherico-subdeprcsscd, about 1 nnu. broad, chcdnnt-broivn, shiuiuff ; mass of si)ores 44 A Monograjyh of the Mi/xogastres. black; spores globose, IG — 18// diameter; episporc blackish violet, smooth. Protoderma ]p%silla, Host., Mon., p. 90. Protodermiwm imsillum, Sacc, Syll., Vol. vii., pt. I, n. 1139. On rotten wood. Germany; Russia; United Status. (Rostafinski's Synonyms.) Zicea pusilla, Sclirad., Nov. Gen., p. 19, t. vi., f. 4 (1797). Pliysarum licea, Fr., S. M., iii., p. 143 (1829). Sub-sect. Cribrariae. Enteridium, Rost. (emended). Aethalium covered with a general, irregularly perforated, membranaceous cortex ; walls of component sporangia with symmetrical perforations, or irregularly broken up to form a capillitium-like structure. Rost., Mon., IX 226; Cooke, Myx. Brit., p. .56; Schroeter, p. 103; Sacc, Syll, Vol. vii., pt. I., p. 409. Very closely allied to the CkctliroptijcMum, which differs chiefly in the more complete absorption of the walls of the sporangia. In E. olivacenm, which most nearly approaches CJatliropty- chiitm in structure, the side walls of the sporangia are not so completely absorbed as in the last-named genus, hence the thickened angles formed by the walls of contiguous cells have broader and more irregular wings. Distrih. Europe ; United States ; W. Australia : species 3. A. Spores Beticulatcd. E)ifcridii(m olivacctt.m,'Rost. (f. 14 — 18). Aethalium very variable in form, flattened or pulvinate, olive ; general cortex papery, pellucid ; walls of the crowded sporangia pellucid, yellowish-brown, perforated; mass of spores olive; spores in clusters of from 5 — 20, single sjwrcs truncate, the free portion convex ami wartcd, rcmaini^ig 2'>ortion smooth, 11 — 15 /x diameter. Enteridium. 4 5 Rost., Mon., p. 227, fig. 5 ; Cooke, Myx. Brit., p. oG, fig. 5 ; Schroeter, p. 103; Sacc, Syll., n. 1399. Eetimlaria ai^iAancda, B. and Br., Ann. Nat. Hist. (188G), t. ii., f. 8. (Specimen in Herb. Berk., named by Rostafinski.) On wood. Britain (Kew, Scarborough, Appin, N. B.) ; Germany ; Denmark ; France ; United States ; W. Australia. Irregularly effused for 1—2 inches, or sometimes in the cracks of wood and then elongated ; colour dark greenish olive, the minute sporangia showing through the thin general cortex, and giving the surface a slightly papillose appearance. The side walls of the sporangia are more or less regularly perforated, the angles where adjacent sporangia meet being permanent and winged with the side portions of the walls that are not absorbed. (Rostafinski's Synonyms.) LycoiKvdon undulimLm, Schum. Saell., 1404 (1803). Enteridium olivaceum, Ehr., f. 1, 5 A.E. (1818). Reticnlaria versicolor, Fr., Obs., ii., 147 (1818). Licogala olivacea, Link, Hb. Betioularia olivacea, Fr., Syst. Myc, iii., 89 (1829). Rcticularia unyidina, Fr., in Fl. Dan., t. 1977, f 2 (1823). Enteridium airum, Preuss., Linnaea, 142 (1851). Beticidaria ajyjylanata, B. and Br., Ann. Nat. Hist. (1866), t. ii., f 3 ; Cke. Hdbk., No. 1098. Zicea olivacea, Fckl. Symb., 338 (1869). Zicca glomulifera, De Bary and Rost, in Alex. (1872). Licaethalium olivaceum, Rost. Vers., p. 4 (1873). LindUadia versicolor, Rost., in Fckl, 68 (1873). Enteridium Rostrupii, Raunk. Aethaha irregular, lengthened, the greatest length is 4 cm., consisting but of one stratum of sporangia, which is 1 mm. thick. Wall of sporangium olive; the lateral wall towards ripening of the spores perforated by large, oval openings. Mass of spores olive; the single spore ratlier colourless. Spores in 4G A Monograph of the Myxogastres. oval hall-shaped clusters of from 5 — 25. Only the convex surface warted, 11 — 12 ix diameter. Enteridium Bostriqni, Rauuk., Myx. Dan., p. 1 OG (in English). On fir-stumps and rotten oak branches — autumn. Denmark. From the above description alone it is difficult to indicate the points of difference between the present species and E, olivacetom. Enteridium macrosporum, Raunk. Aethalia hemispherical, 2 mm. diameter, at first reddish, later olive. Spores in oval ball-shaped clusters of from 5 — 25. Only the convex surface s'pinulose, 12 — 14 /x diameter. Entcridktm macrosporum, Raunk., Myx. Dan., p. 106, t. 2, f. 1. On rotten fir branches. Autumn. Near Copenhagen. This species I have only fo\md once, and then but very few specimens. It seems to be especially different from the other species of Enteridium in the size of the spores and the par- ticularly developed sculpture of their convex surface. (Raunk.) Enteridium Rozeanum, Wing. (f. 108). Aethalium of irregular shape, - globose, ovoid, or rounded- pyramidal, attached to the substratum by a wide base. Vari- able in size, from 5 — 30 mm. in diameter. Cortex and mass of spores ferruginous brown ; occasionally the cortex shining ; sometimes membranaceous, pellucid. Interior structure very variable, the walls of the individual sporangia (which form a capillitium) membranaceous, pellucid, band-like, combined into an all-sided network attached uniformly to all sides of the cortex. The bands have triangular or polygonal expansions at the angles where they join each other. Spores globose, about two-thirds of the surface covered with a delicate, regular, fine-meshed network, the remainder with simple warts or elongated ridges. The warted portion has the outline of a wide spindle lying at the poles of the little sphere. Spores measure 7 5 — 9 ^. Enteridium. 47 Eiitcridiniii Bozcanum, Wing., Proc. Acad. Nat. Sci. Phil., 1889, p. 15G. Beticularia (?) Eozcana, Rost., Mori,, SuppL, no. 179 ; Sacc, Syll., no. 1425. Licea fallax, Pers., in Herb. Schwein. On wood, bark, &c. Britain (Scarboro') ; France; United States. The spaces in the capillitium, wliich represent the individual sporangia of the aethalium, have about the same dimensions in the same plant, but vary considerably, comparing one speci- men with another. The extremes of variation may be ex- pressed by the proportion 1 to 10. Sometimes the bands of the capillitium are very much lacerated longitudinally, and form a filamentous mass, when the characteristic structure of the genus is entirely lost after the dispersal of the spores. Capillitium yellowish under the microscope. The whole of the above is from Mr. Wingate's article, which also contains an interesting historical sketch of the species. The present species appears to be generally distributed in the United States, judging from the specimens in the Kew herbarium sent by various American correspondents; one specimen communicated by Ravenel from Lower Carolina is irregularly circular in outline, three and a half inches in diameter and half an inch thick. The European form, judging from the specimens I have seen, has the sporangial walls very much reduced, and will probably be found in herbaria under Beticularia, where I was at first disposed to place it until con- vinced by Mr. Wingate of its nearer affinity with Enteridium. B. . S'porcs wartcd. Enteridium simulans, Rost. Very similar to B. olivaccum, but the spores are free or loosely aggregated, irregularly spherical, and minutchj wartcd all over, 10 — 12 IX diameter. Rost., Monogr,, Appendix, p. 81 ; Karst., ^Mvc. Fran., iv. p. 12G; Sacc, Syll, n. 1400. 48 A Monograph of the Myxogastres. On decorticated brandies of Almis incana. Finland. Aethalinm variable in form, suborbicular or elongated, convex, pulvinate or depressed, 2 — 5 mm. broad. I have had no opportunity of examining the present species. If the spores are minutely warted all over, the species may be distinct, but clustered spores with warts all over are not common ; and remembering the numerous inaccurate descrip- tions in the works of Karsten and Rostafinski in connection with epispore marking, I am inclined to suspect the autonomy of the present species. Orcadella, Wing. Sporangia with coarse stipes; sporangium wall likewise coarse, but at the top of the sporangium replaced by a delicate membrane which forms a more or less flattened deciduous lid. Orcadella, Wingate, in Proc. Acad. Nat. Sci. Philada., 1889, p. 280. The present genus is considered as the type of a new family of the Myxogastres, described as follows by Mr. Harold Wingate in the article quoted above — " This curious minute plant presents anatomical character- istics which lead me to propose a new family of the Myxo- mycetes, and which, in Dr. Rostafinski's classification, would come imder Order IV., Ancmeae, and after Family 13, Clathro- ptychiaceac. " OTcadcllaccac, fam. no v. " Sporangia without columella or capillitium, and the upper part of the coarse sporangium wall replaced by a delicate membrane having a defined outline. " This family seems to bridge, to a certain extent, the gap between Rostafinski's orders Ancmeae and Hetcrodermcae, as we here have a stage between the uniform wall of Licea, Tuhulioia,, &c., and the lacerate upper wall of some species of Crihraria, where the receptacle is strongly developed and covered with minute granulations. The epispore seems to be absolutely without thickenings even under very high magnification." Orcadella. 40 I am indebted to the kindness of the author for an oppor- tunity of examining authentic specimens, and quite agree as to the validity of the generic characters, and also admit that the new family founded is quite as good as most others at present admitted iu the Myxomycdcs, but I am afraid that if such trivial characters are admitted as sufficient for family characteristics, those left for generic distinctions will be reduced to inappreciable quantities. Compared with allied genera, as Clatltroptijcldnm and Entcridiitm, we find that in the genus under consideration, the ajncal portion of the sporangium is differentiated and event- ually disappears, whereas in the two last-named genera the differentiation and disappearance of the sporangium is confined to certain portions of the lateral wall; or finally, it may be looked upon as a scattered, stipitate Tubulina with a dehiscent apex to the sporangium. Distrih. United States: species 1. Orcadella operculata, Wing. (f. 201). Sporangia stipitate, without columella or capillitium, very variable in form, barrel-shaped, urn-shaped, vase-shaped, jar- shaped, ovoid or nearly globose, or all varieties appearing in one small group ; stipe slightly tapering, erect or bent, blackish, rough and coarse from deposits of plasmodic refuse ; very variable in height. Sporangium wall likewise coarse, blackish, containing deposits of plasmodic refuse, but at the top part of the sporangium replaced by a delicate, yellowish, iridescent, lustrous or vernicose membrane which forms a flat or slightly convex circular deciduous lid, sometimes smooth, sometimes reticulately wrinkled. Mass of spores yellowish. Individual spores almost colourless, globose or irregularly roundish, smooth, 8 — 11 ju, in diameter. Orcadella oj^erciilata, Wing., Proc. Acad. Nat. Sci. Phil., 1889, p. 280, with fig. (Type in Herb. Acad. Nat. Sci. Philada.) On truul^s of living red-oak trees {Oucrcus ruler). United States. 50 A Manor/ i' ft ph of the Mi/jcogastres. I have given the author's own diagnosis of the species. Scattered in cracks of the bark, from -5 — 1 mm. high, some- times a Httle more, stem suberect, sometimes rather flexuous, sporangium very small. Clathroptychium, Rost. Aethalium naked ; sporangia sessile, prismatic or cylindrical, densely crowded on a hypothallus, walls of sporangia persistent at the apex, lateral walls reduced to 5 — 6 slender threads which extend from the hypothallus and support the apical portion. Rost., Mon., p. 225 ; Cooke, Myx. Brit., p. 55 ; Schroeter, p. 1G4- ; Sacc, Syll., p. 405 ; Zopf, p. 137. Rostafinski was the first to point out the true aethalioid nature of Glathroptychiam rugulosum, and in his diagrammatic arrangement, Mon., fig. 80, represents the sporangia as circular in outline, packed close together, and consequently leaving triangular interspaces; each sporangium is represented as having six triangular thickened portions of its wall projecting into the interior of the sporangium. A careful examination of several specimens in various stages of development show that Rostafinski's interpretation is not correct. The sporangia, when young, are cylindrical, with both ends slightly convex, and the walls entire; towards maturity the lateral walls, due to mutual pressure, become flattened, and the sporangia are then hexagonal or rarely pentagonal in transverse section ; at the angles where three contiguous cells meet the walls of the sporangia become thickened, agglutinated together, and persist as the upright threads that support the slightly convex, persistent, cap-like portions of the sporangia. From the above description it will be seen that the triangular upright threads correspond in position to the triangular interspaces in Rostafinski's diagram, and further, that the black triangular thickenings shown in the diagram have no existence in nature. During the period between the formation and maturing of the spores, the fiat lateral walls of the sporangia are dissolved, with the exception of a narrow jagged wing projecting from each aogle of the upright permanent portions. In G. rugulosum these wings are Clath I •optiirh 'mm. 5 1 very rudimentary, but in C. Bcrkcleyi arc often broader than the triangular portion, and recalling to mind the similar struc- ture in Enteridium olivaceum. The triangular upright threads almost invariably show traces of a central cavity, which cor- responds to the intercellular space between the originally cyhndrical sporangia. The persistent apices of the sporangia, when viewed from above under a low power, are seen to consist of very slightly convex hexagons, rarely pentagons, so firmly agglutinated together that the whole can be removed as a continuous membrane ; if this is done, and the membrane then viewed from the under side, after the spores have been cleared away, the stumps of the triangular threads will be seen to originate from the angles where the permanent apices of three sporangia meet ; if this membrane is treated with potassie hydrate, the apices separate from each other as do also the triangular threads ; this separation sometimes takes place in water alone, when the apices with their threads appear as represented in Eostafinski's figures 28 and 29. The above account applies to the typical structure, but both species appear to be very variable. Zopf has figured an aberrent form of G. rugidosurti, haVing the threads supporting the apical persistent portion of the sporangium varying in number, irregu- larly branched and anastomosing. I have met with a similar form, also another variety where the threads are branched and anastomosing and not confined to the margin of the apical portion, but originating from any portion of its under surface, and approximating so closely in structure to certain forms of the genus Pericliacna as to be almost indistinguishable. Distrih. Europe ; Africa ; India ; Ceylon ; Australia ; United States : species 3. Clathroptychium rugulosum, Rost. (f. 25 — 28). Hypothallus well developed, often extending as a whitish membrane for some distance ; sporangia prismatic, apex slightly rounded, and supported by five or six slender triangular per- sistent threads, the fiat face iwinting to the interior of the sporan- gium rugidose ; colour of mass of sporangia reddish, bright •52 A Monof/fffpli of the 3fi/xo(/astres. brown, or umber; mass of spores clear yellow, ochraceous, or tinged with brown, spores globose, very minutdy vcrruculose, 8 — 11 M diameter. Host., Mon., p. 225, figs. 25, 28, 29, 30; Cooke, Myx. Brit., p. 55, figs. 25, 28, 29, 30 ; Scliroeter, p. 104. Licea tcnuissima, B. and Br., Fungi of Ceylon, in Journ. Linn. Soc., Vol. xiv., p. 86 (Hb. B., n. 10858). Zicea cinnaharina, B. and Br., Ceylon Fungi, in Journ. Linn. Soc, Vol. xiv., p. 80 (Herb. Berk., 10854). Ostoricoderma spadiccum, ScliAvein. (specimen so named by Schweinitz in Herb. Berk., 10864). Cornuvia Scliwcinitzii, Berk, in Herb. Eu'sicc. — Ellis, N. Amer. Fung., 335. On dead wood. Britain (Oxford, Leicester, Eltham Park, Scarborough, Glamis, N. B.) ; France ; Germany ; Italy ; Bel- gium ; Hungary ; Algeria ; United States ; Ceylon ; Australia. The crowded sporangia form compact cakes of irregular form, varying from a \ to 1^ in. across, and about 1 mm. thick. The surface is generally chestnut-brown, often with olive shade, and minutely rugulose. The hypothallus often extends con- siderably beyond the sporangia. Differs from C. cinnaha- rinum in the verruculose spores, and from C. Bcrkelcyi in the absence of wings to the triangular supports of the apex of the sporangium. (Rostafinski's Synonyms.) Fuligo phtmhea, Fl. Dan., 1976, f. 1 (1803). Reticularia plumhca, Fr., S. M., iii. 88 (1829). Licea rugulosa, Wallr, Fl. Germ., 2107 (1833). Licca applanata. Berk., Hook Journ. (1845) ; B. and Br., Ann. Nat. Hist., n. 313 ; Cke, Hdbk., n. 1195. Lycogala Icnticulare, D. R. and M., Fl. Alg., 401 (184G). Reticularia leoiticularis, Mort., Herb. Dictydiacfhalium cqyplanatum, Rost., in Fckl., Symb., 2 Nach. 69 (1873). Clathvopt jfch inm. 5 3 Clathroptychium Berkeley!, Mass. (n. sp.). Hypothallus stout, not extending beyond the sporangia, which are densely crowded, forming cakes 1 — 3 cm. across, and 1'5 — 2 mm. thick, very minutely and indistinctly rugulose, brownish-olive ; apices of sporangia almost plane, supported by 5 — 6 triangular, holloio threads, vjhich are strongly winged at the angles, free edges of wings eroded ; mass of spores dirty olive ; spores globose, warted, warts with a tendency to become elon- gated, 10 — 13 fJL diameter. Licca cijlindrica, Fr., Berk, and Br., Fungi of Ceylon, Journ. Linn. Soc, Vol. xiv., p. 86. (Type in Herb. Berk., n. 10857 a.) On wood. Ceylon. The large spores with the warts showing a tendency to become elongated and form ridges, and the strongly-winged threads supporting the apices of the sporangia characterize the present species. Clathroptychium cinnabarinum, Sacc. Sporangia prismatic, broadly diffused, closely crowded, \ — I mm. high, vivid vcriiiilion, above (operculum) blackish- purple, rather shining, sessile, rising erect from a hypothallus ; spores globose, smooth, 9 — 12 ju, diameter, vermilion, guttulate ; threads descending from the operculum, filiform, thickish, fuliginoiis. Sacc, in Michelia, I., p. 545 ; Sacc, Syll., Vol. vii., pt. I., n. 1397. On decayed oak wood. North Italy. Forming flattened heaps, 4 — 6 cm. long. Differs from C lathropt jjchiitm rugulosum in the different colour of the mature plant, the narrow, prismatic sporangia, and smooth, vermilion spores. The vermilion colour and smooth spores suggest the possi- bility of the present being an immature condition of some species. Clathroptychium dissiliens, Hazl. Sporangia forming a rotund or oval cushion, 2 — 5 mm. 54 A Motwr/j-ajyh of the Mi/x<>(faf?tres. diameter, chestnut-brown, pruinose ; mass of spores and sup- porting threads yellowish brown. Dictijdiathaliu.m dissilicns, Hazl., Just, Bot. Jahresb., 1877, p. 156 ; Sacc., Syll, Vol. vii., pt. I, n. 1398. Hazlinsky states the present species, when ripe, dehisces in an elastic manner on the slightest touch, brought about by the sudden expansion of the elaters (= threads supporting apical portion of sporangium) which in(n'ease to three times their previous length. Cribraria. Pers. (emended). Wall of sporangium single, with thickened permanent portions arranged in the form of more or less parallel ribs at the base, branching and anastomosing above to form an irregular net- work, angles of network sometimes dilated and enclosing granules; thin portions of wall entirely disappearing at ma- turity, or the basal portions between the radiating ribs per- sistent and forming an imperforate calyculus ; columella and capillitium absent. Cribraria, Pers. Syn., p. 192 ; Rost., Mon., p. 57 ; Cooke, Myx. Brit, p. 57; Schroeter, p. 104; Zopf, p. 141; Sacc, SylL, Vol. vii., pt. I., p. 412. Hderodidyon, Rost., Mon., p. 231 ; Cooke, Brit. Myx., p. 57 ; Sacc, Syll., p. 411. After a careful comparison of specimens or figures of all the species, with two exceptions, I find it impossible to retain Rostafinski's genus Hderodictyon. In Cribraria we find, as a rule, a calyculus at the base of the sporangium, composed of more or less parallel ribs, with the thinner intermediate portions persistent ; from the irregular margin of the calyculus, the upper network commences, or rather, may be looked upon as a continuation of the ribs of the calyculus; if the upper portion starts from the margin of the calyculus, as an irregular net-work consisting of well-defined nodes united by thin strands, it represents Rostafinski's conception of Cribraria; but if the lower portion of the network consists of more or less parallel, broad ribs, joined laterally by thin strands, and Crlhiutrld. 55 passing into the Cribraria type of network towards the apex, then, according to Rostafinski, it is Hctcrodidyon. There is, in reality, every shade of transition between the two extremes, and G. Blcniaszii closely connects the genus Cribraria in the wider sense, with Dictyilmm. Distrib. Europe; United States; Ceylon: species 19. Sub-Gen. Hctcrodcrma. Calyculus 'present. Network thickened, and forming nodes at the angles. Cribraria elegans, B, and C. Sporangia globose, stipitate, more or less cernuous, p%trj)le, calyculus occupying nearly lower half of sporangial wall, with numerous radiating ribs, and studded with organic purple granules inside ; nodes of network numerous, large, angular or irregularly stellate, containing purple granules, connected by thin bands; stem elongated, expanding downwards, and pass- ing into a small hypothallus, curved near the apex, p)U'^'l>^^ »* spores — mass purple ; spores globose ; minutely warted, 5 — 7 joi diameter. Cribrao'ia elegans, B. and C, Grev., Vol. ii., p. G7 ; Sacc, Syll., n, 1412 ; Rost., Mon., App., p. 82. (Type in Herb. Berk., n. 10873.) On decayed wood. Lower Carolina. Scattered or gregarious, stem variable in length, 2 — 3 mm., sometimes flexuous. There are no free ends in the network of the sporangium. Cribraria piriformis, Schrad. Sporangia 2W^tform> stipitate, erect, p^urplc-brovjn before de- hiscence, then yellowish-brown above, calyculus funnel-shaped, occupying the lower third of sporangium, with radiating ribs, margin eroded, nodes of network large, angular, coloured, with prominent, acute processes, containing granules, united by very slender threads ; stem about equal in length to sporangium, sub- equal, purple-brown, expanding at the base into a hypothallus of the same colour ; spores very pale salmon-colour, globose, smooth, o — 7 fj. diameter. 56 A Manor/ r(iph of the Mjijcona^tres. Crihraria piriformis, Sclirad., Nov. PI. Gen., p. 4, pi. S., f. 4—5 ; Rost., Mon., p. 237, fig. 14; Cooke, Brit. Myx., fig. 14 ; Sacc, Syll., n. 1416. On rotten pine wood. Scotland ; Sweden ; Germany. Gregarious, several often springing from a common hypo- thallus; 2 — 3 mm. high, resembling superficially some forms of C. macrocarpa, but at once distinguished by the large, irregularly stellate, distinct nodes of the network, connected by slender strands. Cribraria macrocarpa. Schrad. (f. 110). Sporangia hroadhj oJwraie, stipitate, erect, upper portion ycUowisk-hroum, calyculus Irownish-jnirplc, funnel-sJiajwd, occu- pying one-third or more of the sporangium; sometimes ^)c?-- f orated toivards the very irregular margin, the projections of which are prolonged as flattened, granulose, coloured, iiTegularly curved bands of varying width, and anastomosing at various points, with numerous lateral projections which are continuous with the projections of adjoining ribs as thin strands bridging the irregular meshes formed by the larger ribs; stem stout, erect, simple, or 2 — 8 more or less grown together, purple- Irown, rugulose, expanded at the base into a spreading hypo- thallus; spores pale yellow, globose, smooth, 7 — 10 ^ diameter. Crihraria iiwcrocar'pa, Schrad., Nov. PI. Gen,, p. 8, t. ii., f. 3—4; Cooke, Myx. Brit., p. 59; Sdcc, Syll., n. 1417; Rost., Mon., p. 238. On decayed wood. Britain (Glamis, N. B.) ; France ; Switzer- land ; Germany ; Sweden. Crowded or scattered, sometimes 2 — 3 more or less combined to form an aethalium. The obovate sporangia and peculiar structure of reticulation characterize the present species, the broad, flattened, primary ribs by irregular bifurcation, combine to form a network of irregularly elongated meshes, which are bridged over by lateral thin strands from the sides of the larger ribs. Well defined distinct nodes connected by thin strands are absent. Criljrari((. 57 (Ilostafinski's Synonyms.) Grihraria viacrocarpa, Sclirad., Nov. Gen., p. 8, t. ii., f. 3 — 4 (1797). TricJda macrocarpa, Poir., Ency., 1. c. viii., no. 80 (1808). Cribraria purpurea, Sclirad. (f. 109), Sporangia globose, stipitate, darh imrplc, calyculus well- developed, with thickened ribs, sometimes irregularly perforated near the dentate margin; nodes of network containing purple granules, small, usually elongated, but very irregular in form, with numerous processes, connecting threads thin, some of the processes are 2)rolongcd into simiilc or branched irregularly bent threads that remain free at the tips, and not connected with other nodes or threads ; stem elongated, flcxuons, purple ; spore- mass dark purple, spores almost colourless, or with faint tinge of rose, globose, smooth, 5 — 7 /x diameter. Cribrao-ia purpurea, Schrad., Nov, PI. Gen., p. 8; Post,, Mon,, p. 233 ; Sacc, Syll, 1408. On decayed pine trunks, Germany; Britain (specimen in Herb. Kew marked, " Herb, Dawson Turner, British ") ; Sweden (specimen collected by Lindblad, and communicated to Berkeley by Fries), Gregarious or scattered, about 1"5 mm, high, stem 3 — 4 times as long as sporangium, usually more or less flexuous. Agrees in colour with C. elcgans, but readily distinguished by the smooth spores and numerous free ends in the permanent network of the sporangium. Distinct from C. macrocarpa in the globose sporangium and long, slender, flexuous stem, Cribraria aurantiaca, Schrad. (f, 104), Sporangia globose, usually more or less cernuous, orange brown, calyculus occupying one-third or more of the sporangial urill, furnished with ribs which pass upwards to form the net- work, nodes colotired, containing gramdcs, large, irregular, united by thin, colourless bands ; stem elongated, attenuated upwards, where it is curved, briglit broivn, expanding at the base into a 58 A Monograph of the Ifyxogasives. small hypothallus ; mass of spores dark orange ; spores globose, smooth, 5 — 7 II diameter. Cribraria aurantiaca, Scliracl, t. i., f. 3 — 4 ; Rost,, Mon., 228, f. 21; Cooke, Myx. Brit., p. 58, fig. 21; Schroet., p. 105; Ilaunk., p. 50 ; Sacc, Syll., 1409. On decaying wood, especially pine. Britain (Carlisle, Aboyne, Linlithgow); Sweden; Germany; Belgium; Hungary. Gregarious or scattered, 2 — 2-5 mm. high ; the nodes of the network of the sporangial wall are numerous, large, and very irregular, usually irregularly stellate, the rays tapering gradu- ally and without much smooth internode, again expanding as a ray of an adjoining node. Spores sometimes minutely verruculuse under a high power. (Rostafinski's Synonyms.) Crihmria aurantiaca, Schrad., t. i., f. 8—4 (1797) ; Fr., S. M., iii. 174; Cke., Hdbk., 1169 ; B. and Br., Ann. Nat. Hist., 1087. Cribraria vulgaris, /3 aurantiaca, Pers., Syn., 194 (1801). Trichia rufcsccns, (3 aurantiaca, Poir, Ency., viii. 81. Cribraria, aurantiaca y sidphurca, Wallr., Fl. Germ. (1888). Cribraria intermedia, Berk., Eng. Fl. v., 318 (1880); Cooke, Hdbk., 1166. Cribraria variabilis, Ficinus Dresd., Fl. 260 (1838). Cribraria tenella, Schrad. Sporangia spherical, generally more or less cernuous, stipi- tate, calyculus occujitying about half the sporangium, brown, shining, often perforated above, margin irregularly eroded, nodes numerous, coloured, containing granules, small, more or less circular in outline, connected by very delicate strands; stem elongated, slender, subequal, often flexuous, brown, with a tinge of purple ; spores globose, almost colourless, smooth, 5 — 7 /x diameter. Cribraria tenella, Schrad., Nov. PI. Gen., p. 6, t. iii., f. 2 — 3 ; Rost., Mon., p. 235, f. 28 ; Cooke, Myx. Brit., lig. 28 ; Schrocter, p. 105; Sacc, Syll, no. 1411. On rotten wood. Germany ; Sweden ; Italy. Crihraria. 59 Gregarious, 1-5 — 2 mm, liigh, resembling superficially C. intricata, but distinguished by the nodes of the network being almost circular in outline, and without the prominent angles from which the delicate strands spring in the last-named species. Cribraria intricata, Schrad. (f. 11 — 13). Sporangia globose, erect, stipitate, ochraccoics hrown, calyculus ocmpying half the sporangmm, margin dentate, nodes of network numerous, containing granules, colowed, irregidarly angular, with several large, jjointed, marginal prqfcetions, connected by very delicate strands, often arranged in more or less jxirallel series; stem erect, attenuated upwards, Irown, with usually a purple tinge ; spores pale, globose, wall thick, smooth, 5 — 7 /u. diameter. Crihraria intrieaia, Schrad., Nov. PI. Gen., p. 7, t. iii., f. 1 ; Rost., Mou., p. 287, fig. 27; Cooke, Brit. Myx., p. 59, I 27; Raunk., p. 50, t. ii., f. 7 ; Sacc, SylL, n. 1415. On rotten wood, especially pine. Britain (Lyndhurst, Wey- bridge, Carlisle, Glamis, N. B.) ; Germany ; Sweden ; United States. Gregarious or crowded, 2*5 — 3 mm. high; distinguished amongst species with a large, hemispherical calyculus, by the large, angular, irregularly stellate nodes of the network being connected by slender bands, which are often arranged in parallel pairs. (Rostafinski's Synonym.) Crihraria intricata, Schrad., Nov. PI. Gen., p. 7, t. iii., f 1 (1797). Cribraria minima, B. and C. Sporangia globose, cernuous, stipitate, dusky ochraceous, calyculus hemisjjherical, margin irregularly toothed, thiclrncd rihs rudimentary; nodes of network numerous, irregularly stellate, coloured, not containing granules; stem elongated, attenuated upwards, brownish; spores globose, almost colour- less, smooth, 5 — 7 // diameter. GO A Monogntph of the Mijxogastres. Co'ihrcma minima, B. and C, Grev., Vol. ii., p. 67. Crihraria viinutissima, Rost., Won., App., p. 31 ; Sacc, Syll., n. 1405 (in part). (Type in Herb. Berk., n. 10874.) On pine planks. Lower Carolina. Scattered, about f mm. high, stem erect, or usually curved above. Perfectly distinct from C. minutissiiiia in the presence of a well-developed calyculus occupying the basal half of the sporangium. Most nearly allied to C. intricata, which is distinct in its larger size, presence of granules in nodes of network and minutely warted spores. Rostafinski has evidently not seen the type specimen of the present species, otherwise he could not jDossibly have given it as a synonym of G. minutissima. Cribraria mirabilis, Mass. Sporangium spherical with a small, irregular calyculus; 2Jcrsistent thickened pm^tions of wall continuing for some distance from the base as inore or less parallel, irregular, flattened ribs, joined laterally by thin threads, towards the apex forming an irreo'ular network with large, irregular nodes; mass of spores obscure fulvous, spores smooth, 7 ju. diameter. Hctcrodictyon mirahile, Rost., Mon., p. 231, fig. 16; Cooke, Myx. Brit., fig. 16 ; Sacc, Syll., no. 1403. Germany. The characteristics of the present species are— the small, irregular calyculus, and the great length of the more or less parallel flattened ribs, which only break up to form a reticu- lation towards the apex of the sporangium. Cribraria Bieniaszii, Mass. Sporangia globose, stipitate, brown, calyculus occupying the lower third of the siiorangium, on the inside furnished with a thickened network, margin dentate, teeth 30 — 40, continuing Hpirards as long, linear, parallel ribs united by very thin hori- zontal strands, p)cissing above into a Cribraria-lihe nctivorh, nodes small with from 3 — 6 angles and concave sides ; stem elongated, becoming attenuated upwards, hnnv)i, expanding into a small Crlbrarla. Gl liypotliallus ; spores clear yellow, vxtll smooth, thin, 5 — 7 jj. diameter. Hctcrodictyon Bieniaszii, Racib., Hedw. 18S0, p. 121. On rotten trunks. Poland. Stem 1'5 — 2-5 mm. high, sporangium 0'8 — 1 mm. diameter. Distinguished from C. mirabilis by the larger sporangium and the greater number of ribs forming the lower part of the network. Cribraria vulgaris, Schrad. Sporangia globose or slightly flattened below, stipitate, caly- culus shallow, ribs very thin, jT'ii-allel, crowded, margin regularly dentate, the very thin threads of the network springing from the teeth of the calyculus, and combining to form a dense net, nodes distinct, hut very small, rarely containing a few granules ; stem slender, elongated, often rather flexuous, Iroivn, with tinge of 2yurple, becoming thicker downwards and expanding into a small hypothallus ; spores very pale, globose, smooth, 5 — 7 fJi- diameter. Cribraria vulgaris, Schrad., Nov. PI. Gen., p. 6, t. i., f. 5.; Rest., Mon., p. 234, f. 26 ; Cke., Brit. Myx., fig. 2G ; Schroeter, p. 105 ; Karst., Myc. Fenn. iv., p. 85 ; Sacc, Syll., no. 1410. Exsicc. Rab., Fung. Eur., 676 ; Fckl., Fung. Rhen., 2100. On decayed wood, especially pine. Germany ; Sweden ; Fin- land. Gregarious, about 1'5 — 2 mm. high. Stem straight or curved above. The features of the present species are — the shallow calyculus with thin, crowded, parallel, or more accurately, radiating ribs, and the very small knots of the network. Cribraria elata, Mass. Sporangia globose, erect or sub-cernuous, stipitate, ccdyculus small, funnel-shcqjcd, radiating ribs well-developed, margin regu- larly dentate, teeth elongated, acute, nodes of network numerous, coloured, irregularly angular, crowded with granules, with several prominent, tapering processes, connected by very slender bands ; stem with the calyculus dark reddish-brown, elongated. 62 A M(mo(jraj)h of the Myxogastres. erect or slightly curved above, becoming thicker downwards and expanding into a small hypothallus ; spores pale yellow brown, globose, smooth^ 7 — 8 \x diameter. On wood and twigs. Ceylon. (Type in Herb. Berk., Kcw, no 10871.) Gregarious, 3 — 3'5 mm. high, superficially resembling Cri- hraria microccnya, but distinguished from this and all other species by the small, funnel-shaped calyculus with the very long, tapering, marginal teeth. The spores sometimes show a tendency to granulation under a high power. * * Network not thickened at the angles. Cribraria microscopica, B. and C Sporangia broadly obovate or globose, stipitate, calyculus occupying almost the lower half of the sporangium, without rib-like thickenings, permanent ribs of upper portion thin, eqiial, without granules, not ioicrctssated at the angles, eomhined to form large, irregular jJolygons of about equal size; stem about equal in length to siwrangium, erect, brownish umber : spores globose, smooth, 6 — 7 /u, diameter. Cribraria microscopiccL, Berk, and Curt., Grev., Vol. ii., p. 67 ; Sacc, Syll., 1406. (Type in Herb. Berk., n. 10175.) On shingles. South Carolina. Scattered, about 1 mm. high, stem about equal to sporangium in length, expanding at the base into a minute hypothallus.. Every part dingy ochraceous. Distinct from G. minima in the absence of thickened nodes in the network of the sporangium. Cribraria tatrica, Racib. Sporangia globo.se, rufous-brown, stipitate, stem obscure rufous, straight; lower part of sporangium persistent, forming a calyculus with the margin irregularly crcnato-dentate ; per- sistent thickenings of upper portion of sporangium cdtout equal, not thickoicd at the nodes ; spores yellow, globose, smooth, 6 — 7 {J- diameter. Cribraria. 63 Crihraria tatrica, Racib., Hedw., 1885, p. 170; Sacc, Syll., n. 1407. On rotten wood. Hungary (Tatra). Sporangia f — 1 mm. diameter, stem 2 mm. long. Cribraria rufa, Rost. (f. 103). Sporangia broadly obovate or sub-globose, stipitate, orange- rufous, calyculus well-developed, occupying one-third or more of the sjwrangkcm, furnished with radiating ribs, which extend above and combine to form a more or less regular net not thickened at the angles ; stem Irown, often with a imr2:)le tinge, elongated, erect, attenuated upwards, expanding at the base into a small hypothallus; spores pale brown, often with a decided jDurple tinge, sviooth, 5 — 7 /u. diameter. Crihraria rufa, Rost., Mon., p. 232, f. 15 ; Cke., Brit. Myx., p. 58, f. 15; Schroeter, p. 105; Sacc, Syll., n. 140-i; Raunk., p. 49, t. ii., f. 5. On wood. Britain (Epping, Carlisle, Linlithgow); Germany; Sweden ; Denmark. Gregarious, 2 — 3 mm. high. (Rostafinski's Synonyms.) Stcmonitis rufa, Roth., Fl. Germ., 548 (1788). Crihxtria rufesccns, Pers. disj)., t. i., f. 5 (171)7). Crihraria fnlva, Schrad., t. i., f. 1 (1797). Crihraria intermedia, Schrad., t. i., f. 2 (1797) ; Berk, and Br., Ann. Nat. Hist., no. 1601. Trichia rufesccns, Poir., Ency., viii., no. 31 (1808). Sub-Gen. Homodcrma. Calyculus cdmnt. * Network thickened and forming nodes cd the angles. Cribraria microcarpa, Rost. (f. 134 — 136). Sporangia globose, small, erect or cernuous, stipitate, ccdi/culus cd)scnt, primary ribs of network radUding from apex of stem as elongcded, hroad bands, anastomosing Icdercdly and forming elongated meshes passing upwards into an irregular network of 64 A ]\[(mo(jraj>h of the Myxogastres. very thin threads connecting large, brown, irregularly stellate knots crowded \Yith granules ; stem slender, very muck elongated, straight or flexuous, erect, or curved above, hro'ivnish intrph ; spores very pale, globose, minutely verruculose, 5 — 7 ju, diameter. Crih-aria microcarpa, Rost., Mon., p. 236; Schroeter, p. 105; Karst., Myc. Fenn., iv., p. 86; Sacc, SylL, no. 1413. Exsicc. — Ellis, North Amer. Fung,, 1398 (as Cribraria tenellet, Schrad.) ; Rav. Fung. Car., 79 (as C. intrieatct). On decayed pine-wood and on sphagnum. Britain (Cleve- don, Bristol) ; Germany ; Sweden ; Finland ; United States. Gregarious, 2'5 — 3 mm. high. Distinguished amongst species without a calyculus by the small sporangium and elongated stem ; also by the elongated, flattened, radiating ribs at the base of the network, thus approaching the type of structure on which Rostafinski's genus Hetcroclictyon was founded, Cribraria splendens, Rost. Sporangia globose, stipitate, erect, yellowish, calyeulns not differentiated, membrane bettveen the thickened portions for a long time 2Jcrsi$tent ; nodes large, angular, containing granules ; stem elongated, slender, sometimes flexuous, pm^i^k-brown ; spores pale yellow, globose, smooth, 5 — 7 /^ diameter. Cribraria splendens, Rost., Mon., p. 236 ; Karst., Myc. Fenn., iv., p. 85; Raunk., Myx. Dan., p. 50, t. ii., f. 6; Sacc, SylL, n. 1414. On rotten pine-wood. Germany ; Finland ; Sweden ; Switzer- land. Scattered, about 2 mm. high, stem about three times as long as sporangium. Distinguished by the very persistent nature of the membrane between the thickened portions. (Rostafinski's Synonyms.) Dictydium splendens, Schrad., Nov. PI. Gen., p. 14, t. iv., f. 5—6 (1797). Grih-aria spilendens, Pers. Syn., p. 191 (1801). Triehia splendens, Poir., Eucy., 1. c. viii., no. 28 (1808). Crihraria. 65 Cribraria dictydioides, Cke. and Balf. Sporangia globose, stipitatc, dingy ochraceous, calyculus absent ; permanent ribs broad and Jiaticned below, anastomosing laterally, filled with granules, passing upwards into numerous elongated or irregularly angtdar, 2^'^^om'inently convex, coloured nodes containing granules, and connected at various i^oints by very thin, colotirless threads; stem elongated, slightly thinner upwards, straight or flexuous, from dirty ochraceous to brown ; spores globose, almost colourless, minutely v&rrucose, 5 — 7 /x diameter, Cribraria dictydioides, Cke. and Balf., Rav. Fung. Amer,, 475. (Type in Herb. Kew.) On wood. Aiken, S. Carolina, and Philadelphia, U. S. Gregarious, about 3 mm. high. The permanent portion of the sporangium closely resembles that of C. argillacca, but in the latter the spores are warted and the sporangium sessile. The present species also agrees with C. microcarpa in the absence of a calyculus, but in the last-named species every part of the permanent portion consists of irregularly stellate nodes connected by thin bands, and the spores are warted. In C. dictydioides, the nodes of the upper portion of the net- work are convex and very prominent. The stem is sometimes branched, or composed of two or three stems more or less completely blended together. Cribraria argillacea, Pers. (f. 105 — 107). Sporangia crowded or scattered, shortly stipitate or sessile, subglobose or broadly pyriform, thin portions of sporamjium very persistent, dirty ochra.ceous, shining, calyculus absent ; thickened portions of sporangium in the form of long, more or less parallel, laterally connected ribs, passing upwards into an irregular network with irregular nodes containing granules ; the internodes have frequently a central, swollen, fusiform, granular portion ; mass of spores dingy ochre ; spores globose, ■minutely warted, 5 — 7 /x diameter. 66 A MonograpJi of the Mijxogastres. Crihraria argillacea, Pers., Obs., i., p. 90 ; Rost., Mon., 239 ; Cooke, Myx. Brit., 59; Sacc, SylL, 1418; Raunk., p. 51, t. ii., f. 8. Exsicc. — Cooke, Fung. Brit, ser. ii., 526. On wood. Britain (Smetbwick, Brandon, Carlisle) ; Germany ; Denmark ; France ; United States. Characterized by the sessile or subsessile sporangia, absence of calyculus, and in the granular swellings of the thickened portions of the capillitium not being confined to the nodes. (Rostafinski's Synonyms.) Stemonitis sphaerocaoya, Schr., Bot. Mag., xii. 20 (1790). Stemonitis argillacea, Pers., in Gmel., 1409 (1796). Crihraria argillacea, Pers., Obs., i. 90 (1796) ; B. and Br,, Ann. Nat. Hist. (1865), no. 1036; Cooke, Hdbk., 1168; Fung. Eritt., ii. 526. Crihraria microincs, Schrad., t. ii., f. 1 — 2 (1797). T7'ichia argillacea, Poir., Ency., viii., no. 32 (1808). ■^ Network not thickened at the angles. Crihraria minutissima, Sz. Sporangia globose, stipitate, calyculus ahsent, permanent ribs thin, forming a more or less uniform reticulation ; angles not at all, or very slightly enlarged, rarely enclosing a few granules ; stem filiform, larger than sporangium, erect or curved above, brownish ; spores globose, almost colourless, minutely warted, 4 — 6 ju, diameter. Crihraria minutissima, Schweinitz, in Syn. Fung. Amer. Bor., no, 2362 ; Rost., Mon., App., p. 31 (Excl. Syn. C. minima, B. and C.) ; Sacc, Syll., 1405 (Excl. Syn. C. minima, B. and C). (Specimen in Hb. B. from Schweinitz.) On decaying pine wood. United States. Scattered, very minute, from "5 — 1 mm. high, recognized at once by the absence of a calyculus and absence of enlarged, Dictydium. 67 angles in the network. Stem from 2 — 3 times as long as sporangium. Dictydium, Sclnad. Wall of sporangium single ; permanent rib-like thickened portions radiating from base to apex and joined by numerous thinner transverse bands ; thinner portions of wall situated between the ribs disappearing at maturity, or persistent at the base and forming an imperforate calyculus ; columella and capillitium absent. Dictydium, Schrad., Nov. Gen. PI., p. 11 ; Rost., Mon., p. 229 ; Cooke, Myx. Brit., p. 57 ; Sacc, Syll., p. 410 ; Zopf, p. 140. The primary, vein-Hke, permanent portions of the sporangium radiate from the base and converge at the apex without becoming much broken up or distorted, although there is usually an indication of a reticulate arrangement near the apex ; these primary ribs are united laterally by straight, thinner portions, hence the arrangement of the skeleton is into more or less regular rectangular meshes. Distrih. Europe; N.America; S.America; Ceylon; Australia : species 2. Dictydium cernuum, Nees. (f. 183). Sporangium globose, cernuous, more or less umbilicate at the apex, reddish brown ; stem elongated, slightly thinner upwards, with a sharp curve near the apex, above which the stem is hyaline and shrivelled at maturity ; below the curve dark brown, opaque, smooth, passing into a small hypothallus ; calyculus small ; mass of spores purple brown ; spores globose, smooth, 5 — 7 fj. diameter. 'Dictydium cernutcvi, Nees., Syst., f. 117; Rost., Mon., p. 229, figs. 17, 19, 22; Cooke. Myx. Brit., p. 57, figs. 17, 19, 22; Sacc, Syll., no. 1401; Schroeter, p. 106; Raunk., p. 51, Tab. ii. fig. 4. Dictydium cernuum, var. laxum, B. and C, Grev. 1873, p. 07. J^'xsicc. — Fuckel, Fung. Rhen., 1445 and 2099 ; Syd., Myc. 68 A Monograph of the Myxogastres. March., 164; Rab., Fung. Eur., 2519; Jack, Leiner. &c., 422 ; Rav., Fung. Car., 78 ; Rav., Fung. Amer., 476 ; Ellis, N. Amer. Fung., 1122. On decaying wood, especially pine. Britain (Milton, Norths ; Kew, Suffolk, Scarboro', Carlisle, Appin, N. B.) ; Germany ; Sweden ; Finland ; Belgium ; Denmark ; Hungary ; United States ; S. America ; Ceylon. Gregarious, 1 — 1"5 mm. high; the stem varies from 2 — 3 times as long as the sporangium ; the lower three-fourths is erect, blackish-brown, and opaque ; at the top of this erect portion there is a sharp curve, above which the pendulous portion of the stem consists, at maturity, of a colourless, shrivelled membrane, from which the sporangium hangs sus- pended, and easily moved by wind or insects ; in all probability this peculiar structure of the stem is connected with spore- dissemination. The inner surface of the calyculus and the ribs are furnished with numerous dark brown granular masses of organic matter. (Rostafinski's Synonyms.) Mtccor cancdlatus, Batsch, f. 232 (1789). Stemonitis cancdlata, Gmel, Syst., 1468 (1791). SjjJiaerocmytos frichioides, Bull, p. 124 (1791). Crihraria cermia, Pers., Obs., i. 91 (1796). Didydium umhilicahcm, Schrad., t. iv., f. 1 (1797) ; Fr., S. M., iii. 165 ; Berk., Eng. Fl., v. p. 818 ; Cooke, Hdbk., 1165, fig. 134. Didi/dium amhiguum, Schrad., t. iv., f. 2 (1797). Tridiia ccrnua, Poir., Ency., viii., no. 25 (1808). Didydium cernuttm, Nees., f. 117 (1816). Didydium tridiioidcs, Chev., Fl. Par., 327 (1827) ; Corda Ic, v., f. 36. Dictydium venosum, Schrad. Sporangium globose, cernuous, brownish-yellow owing to the spore-mass showing through the translucent wall, which is Dictydiam. 69 almost colourless when empty and furnished with 9 — 12 darker coloured ribs giving off lateral, free, not anadoiiiosiiig branches ; stem elongated, flexuous, brownish. Dictydium venosum, Schrader, Nov. Gen., p. 14, pi. iii., fig. 6; Sacc, Syll., v. 7, pt. L, no. 1412 ? On rotten prostrate trunks of pine. Germany. It is doubtful whether the species described by Berlese in Sacc, Syll., 1. c, is the species intended by Schrader. COLUMELLIFERAE. The most pronounced features of the present section are the presence of a central columella, which in most instances is a direct continuation of the stem within the peridium, and the absence of lime from every part. The two sub-sections as defined appear distinct, but this, as usual, is only true of the two poles, and curiously enough, transitions have been met with where they would have been least expected. In the Lamprodcrmeae, characterized by the capillitium springing from the apex of the columella, we find this character fairly satis- factory in the typical genus Lamprodxrriia, and yet more rigidly carried out in Enerthen&ma, where the copious capillitium originates entirely from a peltate disc springing from the apex of the columella after the latter has passed quite through the sporangium ; yet we find in the genus Anct/roj^horus all the characteristics presented by the genus Encrthencma, plus a few scattered branches springing from various points of the columella throughout its length, in addition to the main bulk of the capillitium which originates from the apical disc as in Encr- thenema. The branches of the capillitium springing from the columella are again broken ujd into smaller and smaller branch- lets, which usually combine to form an irregular network. The sporangial wall is always very thin and blackish in colour and remarkable for its brilliant metallic tints, and also for its ephemeral nature, disappearing entirely or in part as the spores approach maturity, or as in the genus Orthotrichia being absent 70 A Monograph of the Myxogastres. from the earliest condition. In Lamfroderma violaceiim the columella is filled with large spherical, or from mutual pressure, polyhedral cells, a character repeated in the stem of several species of Arcyria. The three genera Amaurochaete, Brefeldia, and Retiadaria are only known in the aethalioid condition, and superficially appear quite isolated from the remaining genera, but structurally are nothing more than confluent forms of such genera as Stemonitis and LamjJroderma. Comatricha confiucns is a connecting link between the two extremes. According to the preconceived idea that the presence of a columella is of sectional importance, the genus Si'plwptychium belongs to the present division, nevertheless its affinity with the Fcritricheae is obvious. The following arrangement indicates the affinities of the genera included in the Colw/ielliferac, and the points of contact with other sections. Stemonitis. Ancyrophorus. Brefeldia.—— Rodajinskia. Amaurochaete. I Reticularia. -Enerthenema. I Lamproderma.- Echiiiostelium. I Orthotricha-. I Raciborskia. X — H ° I H !3 Siphoptychiitm. TURHMXA. I PERITRICHEAE. Stewouitae. 71 ANALYSIS OF THE GENERA. Sub-sect. Stemonitae. A. S2)0')'angia free. Stemonitis. Capillitium threads springing from all sides of an elongated columella. B. Sjiorangia forming an acthalmm. Si23ho2>tycMum. Sporangia straight, parallel, each containing a central columella. Amaueochaete. a large, pulvinate aethalium, bifurcating branches of capillitium connected for some distance by a thin membrane. Brefeldia. Aethalium large, threads of capillitium with clusters of vesicular bodies. Rostafinskia. Threads of Lnver portion of capillitium septate. Reticularia. Aethalium pulvinate ; capillitium threads forming an irregular network without vesicles or flat mem- branous expansions. Sub-sect. Lamprodermae. Enerthenema. Threads of capillitium springing from a flattened disc at the apex of the columella. Ancyroijhorus. Threads of capillitium springing from a flattened disc at the apex of the columella, and also from the columella throughout its length. Lamproderma. Threads of capillitium springing from the abrupt apex of the short columella. Echinostdmni. Columella absent ; capillitium springing from apex of stem. Bacihorslda. Wall of sporangium absent, main branches of capillitium springing from apex of a short columella, becoming branched, branches forming a network. Orthotricha. Sporaugial wall absent from the first. 72 A Mojiograjfli of the Mijxiyjastres. Sub-sect. I. Stemonitae. Stemonitis, Gled. Sporangia cylindrical, ovate, or globose, stipitate ; stem con- tinuing into the sporangium as a columella of greater or less length, and giving origin to numerous branches throughout its length, which combine to form a dense, irregular network ; wall of sporangium thin, soon disappearing, often with metallic tints. Stemonitis, Gleditsch, Meth., p. 141 ; Rest., Mon., p. 193 (in part); Cke., Myx. Brit., p. 40 (in part); Sacc, Syll., vii., pt. I., p. 397. Comatricha, Preuss. in Sturm's Deutschl. (in part) ; , Rost., Mon., p. 197 (in part) ; Cke., Myx. Brit., p. 47 (in part) ; Sacc, Syll., vii., p. 894. The genus Comatricha was distinguished according to Preuss by the globose or shortly elliptical sporangium from the species of Stemonitis, a character shown by Rostafinski to be untenable, hence the last-named author used characters suisposed to be presented by the capillitium for the separation of the two genera. In Comatricha the meshes of the network are arcuate or curved, and not attached to the wall of the sporangium, by short branches, whereas in Stemonitis the threads are not arcuate at the surface, and the capillitium is attached to the wall in numerous places by short branches. These characters, not being constant, and furthermore overlapping in the two genera, cannot be considered as of generic value, neither in my opinion would they be so even if constant. Distrib. Scattered over temperate and tropical regions. Species 23. A. Spores Uacldsh hroton in the mass. § Spores wartccl. Stemonitis fusca, Rost. (figs. 155 — 157 and 1G2— 104). Densely grcyarioiLS, springing from a vxll-dccelopcd, broadly Steiiumitis. 73 spreading hyiiothallus ; sporangia cylindrical, obtuse or attenu- ated at the ajDcx, wall thin, evanescent ; blackish with metallic tints ; stem slender, equal, shorter than the sporangium, blackish, shining ; columella reaching nearly to apex of sporangium ; capillitium dense, threads blackish, tapering, combined into a dense, irregular net, the periplural meshes smaller or equal to the S2Jorcs in diameter : spores in the mass hlacJc, luith violet tinge, dingy violet by transmitted light, globose, varying from dis- tinctly verrucose to almost smooth, size also variable, from 5 — 11 [JL diameter. Stemonitis fusca, Rost., Mon., p. 193, fig. 40; Cke., Myx. Brit., p. 4G, fig. 40; Sacc, Syll, n. 1362. Exdcc.—^owm., Fung. Sel. Gall. (Rel. Moug.); Ellis, N. Amer. Fung., 1119; Fckl, Fung. Rhen., 1149; Holl, Schmidt, and Kunze, Deutschlands Schwamme, n. cix. ; Cke., Fung. Brit., Ed. ii., n. 522 ; Fung. Cub. Wrightiani, n. 538 ; Roum., Fung. Gall, Cent. xiv. (number indistinct). On rotten wood, &c. Britain (Bournemouth, Kew, Ilfra- combe, Epping ; Bulmer, Yorks ; Carlisle, Aboyne) ; Europe ; N.America; Cuba; Venezuela; Natal; Mauritius; Himalayas; Ceylon ; Victoria ; Tasmania. From 5 — 14 mm. high. The only large British species with a small meshed capillitium and spores blackish violet in the mass. (Rostafinski's Synonyms.) Lycoperdon capite cylindracca, Rupp. Jenn., 304 (1718). Clathroidasto'um ohscuriim, Mich., t. 94, f. 1 (1729). Emlolus nigcrrimus, Hall, t. i., f. 1 (1742). Claihonis nudus. Linn., Fl. Suec, 1263 (1745). Stemonitis I, Gled., Meth., 141 (1753). Emholus, Hall, 2137, t. 48, f. 1 (1768). Tubulifcra cremor, Fl. Dan., t. 659, f. 1 (1777). Tremella typhina, Willd. Fl. Ber., 420 (1787). Mucor araneosus, Jacq. Misc., t, 15 (1778). Stemonitis ftcsca, Roth. Mag. Bot, p. 26 (1782); Elir. Ber., f. 5 ; Grev., S. C. Fl., t. 170 ; Corda Ic, ii., f 87 ; Eng. Fl., v., 317; Cooke, Hdbk., 1155, fig. 132, Fung Britt., ii, 522. 74 A Monograpli of the Myxogastres. Trichia mida, With. Arr., iv., 477 (1792). Stemonitis fascicnlata, Pers., Syn., 187 (1801). Stcmonitis tyj^hoides, Auct. Stemonitis maxima, Sz. Sporangia crowded, stipitate, springing from a well-developed hypothallus; sporangia cylindrical; wall blackish, reflecting metallic tints, evanescent; columella reaching almost to the apex of the sporangium; capillitium dense, branches often arcuate, sometimes more or less irregularly dilated, especially at the nodes, peripheral meshes nutch larger than the diameter of the spores; mass of spores hlacJdsh brown; spores globose, very minutely verrucnlose, 5 — 10 /a diameter; stem filiform, black, shining, shorter than the sporangium. Stemonitis maxima, Schweinitz, Syn. Fung. Amer., n. 2349 (specimen from Schweinitz in Herb. Berk., Kew, n. 10835). Stemonitis splc7ulens, E,ost., Mon., p. 195 ; Sacc, Syll., vii. 1, n. 1364. (Authentic specimen named by Rostafinski.) On wood, &c. Russia ; Finland ; France ; Ceylon ; Mauritius ; Cuba ; United States ; Australia ; Cape of Good Hope. A fine large species, distinguished amongst the dark-spored species by the large peripheral meshes of the capillitium. Lumps of organic matter are sometimes met with in consider- able quantities on the threads of the capillitium ; 1'5 — 3 cm. high. Stemonitis typhina, Mass. Gregarious, but not usually crowded, springing from a delicate hypothallus ; sporangia cylindrical, ohtuse a.t loth ends, wall very thin, brownish, disappearing; columella reaching to the apex, capillitium dense, threads originating at numerous points from the columella, dingy brown, copiously branched and form- ing a dense, irregular network, thickest at the point of origin and tapering towards the perijDhery where they are very slender, and form a network of irregular pattern, with the meshes usually not larger than the diameter of the sjmres ; stem erect, slender, cylindrical, usually rather shorter than the sporangium, blackish ; Stemonitis. 75 mass of spores brotrnish-UaeJc, lilac-brown by transmitted light, globose, very minutely asjJo-atc, 5 — 8 /x diameter. Rostafinski gives the following forms, but it is not unusual to meet with both in the same cluster. a. Genuina. — Sporangium with stem 2^ — 4 mm. high. /3. Pumila. — Sporangium with stem 2 mm. high. Comatricha typliina, Rest., Mon., p. 198, figs. 46, 47; Cke., Myx. Brit., p. 47, figs. 46, 47; Sacc, Syll., vii., n. 1355. On rotten wood. Britain (Lyndhurst, Kew, Scarboro', Car- lisle, Aboyne, N. B.) ; Europe ; U. States ; Cuba ; Ceylon ; New Zealand. From 4 — 7 mm. high. Closely allied to G. 2)ulchella, but distinguished by the dark brown colour of the spores in the mass. (Rostafinski's Synonyms.) Glathwidastrum ohsmrum, Mich., t. 94, f. 2 (1729). Muco7' capiUdo fnsco, Scop., Fl. Carn., 66 (1760). Mucor Stemonitis, Scop., Fl. Carn., 493 (1772); Schpf., t. 296. Emhohts lacteus, Jacq., Muse. 1, t. 6 (1778). Clathrus nudtos, Fl. Dan., t. 755 (1782). Clathrus fertnsus, Batsch, f. 176 (1753). Stemonitis typhina, Roth. Fl. Germ., 1, 547 (1788); Pers., Obs., 1, 57. Stemonitis filicina, Schrk., Fl. Bav., 1782 (1789). Tricliia typhoides, Bull., t. 477, f. 2 (1791). Stemonitis typhoides, D. C, Fl. Fr., ii., 257 (1805); Cooke, Hdbk., n. 1157. Stemonitis hicolor. Fries, Herb. Stemonitis leiicopoda, Fr. Gast., 16 (1817). Stemonitis pumila, Corda Ic, v., p. 37 (1842). Stemonitis macrosperma, Mass. Sporangia naked, stipitate ; stem continued into the sporan- gium as a columella, reaching nearly to the apex, threads of capillitium passing from the columella to the wall and pro- 7(j A Monograph of the Myxogastres. ducing very much thinner branchlets, which are combined amongst tliemselves at all points to form a rather loose net- work, the, -pcrvplieral mesJies of the netwoQ'k formed of combined^ arcuate threads, and slightly smaller than the rest, especially towards the base of the cajnllitium ; spores 9 — 12 /x diameter, pale violet, vcrrucidose. Stemonitis macrosperma, Racib., Myx. Crac, p. 8 ; Sacc, SylL, vii„ 1, n. 1361. Var. ohovata. Racib., sporangia obovate, 0'5 — 0'75 mm. broad. With the type. Var. oUonga. Racib., sporangia oblong, 0'75 — 1 mm. long, 0'3 — 0'5 mm. broad. With the type. Poland. Stemonitis Suksdorfii, Mass. Gregarious, stipitate, sporangia cylindrical, very obtuse at both ends, wall iridescent, evanescent ; stem blackish-brown, shiuing, about equal in length to sporangium, tapering upwards, rather stout, becoming dilated at the base into a hypothallus ; spores in the mass hlackish-'jiurple. Columella rather thick, sub-equal, blackish, Visually becoming abruptly divided into branches at some distance below the apex of the sporangium; sometimes more or less flattened at the apex, and with a tendency to become discoid ; capillitium exceedingly dense, dark, main branches originating from the columella; stout, taper- ing, branches arcuate, combined to form an irregular, small- meshed network ; spores bright lilac, minutely loarted, 7 — 8 ju, diameter. Goiiiatricha Sahsdorfii, Ellis and Evorh. On wood, bark, &c. United States. (Described from a portion of type specimen communicated by Wingate.) From 5 — 6 mm. high. Stemonitis affinis, Mass. Sporangia cylindrical, obtuse, up to 2 mm, high ; stem^ from, Us shortness scarcely distinct : columella reaching almost to apex Stemonitis. 77 of the sporangium, then dividing into 2 — 5 branches ; threads of capillitiuni much more rigid at the base than in the ulterior, very loose network, nodes often triangular, ultimate branches arcuate, transversely joined, some free and excurrent; spores pale violet, warted, 5 — 6 /u diameter. Comatriclia affinis, Rost., Mon., p. 202 ; Sacc, Syll, vii., 1 n. 1359. Germany, Stemonitis confluens, Cke. and Ellis, Forming compact tufts about 2 mm. high, from 8 — 15 ; individuals spring from a firm, spreading, venulose hypothallus ; stems usually simple and distinct, dark brown, wrinkled to- wards the base, about -5 mm. high, then hrcaking up into from 2 — 4 main tranches or columellas which anastomose later- ally tcith those of other individuals forming an irrcg^dar plasmodiocarp, covered with a blackish-violet, evanescent, common cortex; mass of spores black with violet tinge; the main branches springing from the columellas form a dense network, threads arcuate, many of the meshes closed by a thin membrane ; spores globose, dingy violet, minutely warted, 7 — 8 ^ diameter. St&monitis confluens, Cke. and Ellis, Grev., vol. v., p. 51. Comatricha confluens, C. and E,, Myx., U. States, p. 896. On oak bark. United States. A very interesting species, and highly instructive in showing how the fundamental simple form becomes modified by con- fluence. The general mass of the plasmodiocarp is sometimes spherico-depressed, and even at others the columella of one individual of the colony remains simple and projects above the mass as a finger-like projection; in others all the individuals are more or less free at the tips. When the wall and spores are completely blown away, the resemblance of the capillitiuni to that of Heticularia is striking. 78 A Monograph of the Mi/xogaatres. § § Sjjwes smooth. Stemonitis atra, Mass. (n. sp.)- Scattered ; sporangia cylindric-ovate, black, slightly iridescent ; stem twice as long as sporangium, thin, equal, dull, black, expanding at the base into a minute, circular hypothallus ; mass of spores hroion with a lilac tinge ; columella as thick as the stem, equal, reaching to apex of sporangium ; main branches scattered, remaining simple for a short distance then branching and forming a network, the meshes becoming smaller towards the periphery where they equal the spores in diameter ; all the tranches of the network of equal thickness, very stout. So — 4 fx diameter; dark brown; spores globose, dingy lilac-brown, s?/ioo^/i, 10 /x diameter. On wood. New Zealand. Remarkable in the scattered habit, 6 — 8 mm. high. Readily distinguished by the very thick threads of the capillitium. The columella sometimes breaks up into branches a short distance below the apex of the sporangium. Stemonitis acuminata, Mass. (n. sp.). Gregarious, or often densely crowded on a well-developed, iridescent hypothallus ; sporangia cylindrical, elongato-acicminate at the a'pcx ; wall blackish-purple, iridescent, disappearing; stem equal to or shorter than sporangium, filiform, slightly attenuated upwards; blackish, shining; mass of spores black with purple tinge ; capillitium continuing to the apex of the sporangium, primary branches scattered, short, sui)2^orting a network of irregularly polygonal meshes which are large and everywhere of equal size, 40 — 50 n in diameter ; spores globose, smooth, 13 — 14 /x diameter. On rotten wood. Cape of Good Hope (Table Mountain). Agreeing with Stemonitis Morgani in the large-meshed capillitium, but distinguished by the acuminate apex of the sporangium and the larger spores ; 1"5 cm. high. Stemonitis. 79 Stemonitis Bauerlinii, Mass. (n. sp.). Densely gregarious on a well-developed, firm hypothallus ; sporangia elongato-fusiform, wall blackish, iridescent, very evanescent, mass of s^wres Uach with a violet tinge ; stem erect, black, shining, slightly tapering from the base; columella ex- tending to apex of sporangium ; fiexuous above ; capillitium of lotocr half of sporangium of large, equal meshes which are peri- pheral only and supported on distant, thin, simple or rarely divided branches springing from the columella, the main branches heeome more and more numerous, thicker, and irregulctr towards the apex of the sporangium, and often form irregular, flattened expansions; the hunches forming the net are also much thicker, and the meshes irregular, and not all p)eripheral as towards the base of the S2wrangium; many of the branches of the nettvork towards the apex with short free ends; spores globose, smooth, 5—6 fi diameter. On wood. New Guinea (Strickland River). From 1-5—2 cm. high. Distinguished by the capillitium being very scanty and the network being entirely peripheral below, becoming very dense towards the apex. Stemonitis laxa, Mass. Sporangia spherical, scarcely -5 mm. diameter; stem rigid, black, not attaining to 1 mm. in length ; columella alm^ost reaching to apex of sporangium, and then dividing into 2—3 branches; threads of the capillitium of equal width throughout, very rigid, forming a regular, lax network, ultimate branches arcuate, connected by transverse bands, some remaining free and furcately divided; spores smooth, lilac, 9—11 /a diameter. Comatricha laxa, Rost, Mon., p. 201; Sacc. SvH vii 1 no. 1358. ' Germany. Stemonitis fluminensis, Speg. Hypothallus very thin, broadly effused, mucedinous, black, rather shining; stem erect, rather rigid, shining, black, 0-5- 1 80 A Monogrnpli of the Miixogaatres. mm. long, 0"7 — 0'6 mm. thick ; smooth when moist, rugulose when dry, sub-contorted, extended into the sporangium as a cokimella, not reaching to the apex; sporangium cylindrical, rarely sub-clavate, rounded at both ends, 0*8 — 1'2 mm. long, 0"3 — 0"2 mm. thick, black, ajyaqiic, wall persistent fm^ a long time; capillitium springing from the columella, forming a rather dense network, the swperjicial meshes equal to or tioice the diameter of the spores, with uncinate free tips ; spores globose or sub-elliptical, 5 — 8 /x diameter, perfectly smooth, smoky brown. Stemonitis Jtuminensis, Spcg., Fung. Arg., N. 261 ; Sacc, Syll., vii., 1, n. 1366. On bark, moss, &c. Brazil. Stemonitis subcaespitosa, Mass, Stems subcaespitose or loosely clustered, thickened at the base, about half the length of the sporangia, extending through the capillitium as a columella; peridia ovate-oblong, obtuse, fugacious ; capillitium growing from the columella, reticulately connected and also forming a superficial net with coarse meshes, blackish ; spores globose, even, (smooth ?), blackish-brown, •0004— -00045 in. in diameter. Comatricha sulcacspitosa, Peck, 43rd Report, New York State, Mus. Nat. Hist. (1890), p. 25, PI. iii., figs. 6—9. Decorticated wood of hemlock. Tsuga Canadensis. Sandlake. July. This species resembles Stemonitis fusca in colour. In size it approaches Comatricha typhina. Its capillitium is variously connected, and appears to combine the reticulation of Comatricha and Stemonitis, but on account of the network not being wholly parallel to the walls of the peridium, it is placed in Comatricha. The plants are mostly connected in small groups or loose clusters of two to ten individuals. Its coarser meshes and larger spores distinguish it from Comatricha typhina. (Peck, 1. c). Stemonitis aequalis, Mass. Gregarious or loosely clustered, about three lines high, arising from a thin hypothallus ; sporangia cylindrical, obtuse, fugacious, Stenioint/'s: 81 ■wholly falling away; capillitium brown or blackish-brown, forming an intricate network ; stem slender, smooth, black, penetrating the capillitium as a columella, and extending nearly or quite to the apex, the free portion about equal in length to one-half the altitude of the entire plant; spores globose, smooth, violet-black, '0003' — '00035' in diameter ( = about 8-9 ,.). ComatricJia aeqiicdis, Peck, 31st Report of State Mus., N. Y., p. 42; Sacc, Syll., n. 1300. Decaying wood. Catskill Mountains, Sept. In colour this species is almost exactly like Stemordtis fusca, from which its more lax habit, proportionally longer stem and different capillitium separate it. The larger size, both of the plant itself and of the spores, will separate it from Comatricha tifpliina. The length of the stem and of the capillitium are nearly equal, hence the specific name. (Peck.) Stemonitis tenerrima, B. and C. Gregarious ; sporangia cylindrical, hasc and apex very obtuse, wall thin, blackish-purple, iridescent, evanescent ; stem about equal in length to sporangium, filiform, blackish, sub-equal ; columella reaching nearly or quite to apex of sporangium ; primary branches short, breaking up into a network, the meshes becoming smaller towards the pcrijjhery where they equal the spores in diameter, the threads taper towards the periphery; spores in the mass black with violet tinge, globose, smooth, 13 — 14 IX diameter. Stemonitis teneri'ima. Berk, and Curt., Grev., vol. ii., p. 69. (Type in Herb. Berk., Kew, n. 10831.) On wood. Lower Carolina. Distinguished by the small size 3'5 — 4 mm., cylindrical sporangium, and the large, smooth spores. Whether the present species is identical with "Stemonitis tenerrima Curtis in Sill., Journ., 1. c, p. 349 (1848)," as quoted by Rostafinski, Mou., Append., p. 27, I have no means of deteiniiuiug. 82 i Monof/raj)/f of the Myxogastres. Stemonitis Friesiana, De Bary. Scattered or gregarious ; sporangia glohosc or aliorthj elliptical, wall very thin, disappearing, whitish with a silvery sheen or purjDle black ; stem hlacJc, shining, elongated, tapering ujnvards, expanding at the base into a small, circular, irregularly ribbed or latticed hypothallus ; columella reaching cdiout two-thirds the height of the sjjorangium, then breaking up into several branches, threads of capillitiuni dark brown, arcuate, forming a dense net; spores in the mass blackish, violet-brown by transmitted light, globose, smooth, 8 — 10 /x diameter. Stemonitis Friesiana, De Bary, in Rab. F. Eng., n. 568. Comatricha Friesiana, Rost., Mon., p. 199, figs. 51 and 56 ; Cke., Myx. Brit., p. 48, figs. 51 and 56 ; Sacc, Syll, vii., 1356. Exsicc. — Fuckel, Fung. Rhen., n. 1446 (as Stemonitis ovata). On wood, branches, leaves, &c. Britain (Lyndhurst, Kew, Hereford, York, Carlisle, Aboyne, N. B.) ; Europe ; U. States ; Ceylon ; Tasmania. From 3 — 5 mm. high. The threads of the capillitiuni are of nearly equal thickness throughout, and sometimes studded with amorphous lumps of organic matter. Distinguished by the long, tapering stem, and small globose, or shortly ovate sporangium. (Rostafinski's Synonyms.) Mucor emholns, Linn. Sp., 1185 (1753). Lycogala, Hall, 2146? (1768). Stemonitis reticulata, Trent, p. 223? (1797). Stemonitis iiigra, Pers., Gmel. Sys., 1467 (1791). Stemonitis atrofusca, Pers., Disp., 11 (1797). Stemonitis atrofusca, ^ nigra, Pers., Disp., 54 (1797). Stemonitis ovata, Pers., Syn., 189 (1801); Berk., Eng. FL, v., p. 317; Cooke, Hdbk., no. 1158. Trichia mucoo^iformis, Schum., Saell., 1469 (1803). Stemonitis violacea, Schum., SaelL, 1491 (1803). Stemonitis nigra, Schum., Saell., 1493 (1803). Stemonitis glohosa, Schum., Saell., 1494 (1803). Stemonilis. 83 Trichia alba, Sow., t. 259 (1818). Siemonifds obtusata, Fr. Sym. Gast., 17 (1818); Eng. Fl., v., p. 317; Cooke, Hdbk., no. 1150. Comatricha obtusata, Preiiss., Sturm. (1851). Comatricha alba, Preuss. in Sturm (1851). Stcmoiiitis Friesiana, De Bary (1870) ; Rabh., Fung. Eur., no. 568. § § § Spores reticulated. Stemonitis longa, Mass. Sporangia gregarious, slender, cylindrical, straight, or usually fiexuouSy very long; wall evanescent; stem filiform, slender, rather short, blackish ; columella, capillitium, and mass of spores brotonisli-violet ; columella extending nearly to apex of sporan- gium, tapering upwards; capillitium springing from the colu- mella at numerous points throughout its length, main branches soon branching, axils usually rounded, often occupied for some distance by a thin membrane, sometimes connected laterally, branches towards the periphery much branched in a penicillatc manner, ti2)s free ; spores globose, epispore very minutely reticu- lated, 7 — 8 /x diameter. Comatricha longa. Peck, 43rd Report of New York State Mus., p. 24, pi. 3, f. 1—5. On wood, bark, &c. United States. Fasciculate, capillitium often curved or twisted after the wall has disappeared, 3 — 6 cm. long. Stemonitis dictyospora, Rest. (fig. 112). Sporangia gregarious, cylindrical, both ends obtuse, wall silvery, soon disappearing, stem shorter than sporangium, rather thick, equal, blade, springing from a well-developed hypothallus ; columella reaching nearly to apex of sporangium, capillitium dense, threads like the columella blackish, combined to form a network having the peripheral meshes not larger t1ian the diameter of the sjiores ; spores in the mass blacldsh-violet, globose, epispore with thin ridges forming a polygonal network, 1 2 — 1 5 ii diameter. 84 A Mo7ioffraph of the Myxogastres. Stcmonitis didi/osjwra, Rost., Mon., p. 195 ; Sacc, Syll., vii., 1, 1803. On palm leaves. Venezuela ; Amazon Valley ; United States. The type specimen on wliich Rostafinski founded the species is in the Berkeley herbarium, Kew, and had evidently been considered as distinct as the MS. name of " S. trechispora, B. and C," is on the paper in Berkeley's writing. It seems a pity that Rostafinski could not accept this name. Distinguished at once by the silvery white sporangial wall, very small peripheral meshes of the capillitium, and reticulated spores. B. Spores brown or ferruginous in the mass. § Sjwrcs smooth. Stemonitis Carlylei, Mass. (n. sp.) (figs. 158, 159). Growing in small scattered tufts, sporangia cylindrical, apex obtuse, wall very thin, evanescent except the apical portion, which remains like a cap, dark, with purple or dark blue tints ; stem short, dark, expanding into a tough, brown, common hypo- thallus ; mass of spores dull oraoigc-browii ; columella attenu- ated upwards, disappearing just below the apex ; capillitium dense, originating in numerous stout branches from the colu-' mella, which soon break up into thinner branches that anasto- mose to form a dense, irregular net, threads arcuate, peripheral meshes large, at least tivice the diameter of the spores, and bearing numerous free tips which are at first attached to the wall, brown, becoming paler towards the periphery; spores globose, pale, clear orange-brown, smooth, 11 — 15 fx diameter. On wood. Carlisle. Growing in small scattered tufts of from 8 — 14 plants, 3"5 — 4 mm. high, the sporangia are cylindrical with abruptly rounded ends, or rarely slightly attenuated below, and passing into the stem, which is not usually a quarter the length of the sporangium. At once distinguished from Stemonitis ferruginca, the remaining bright-spored species by the large peripheral meshes of the capillitium, with numerous free tips, and the Stemo7iitu. 85 larger spores. The spores sometimes sliow indistinct iudicatious of warting under 1200 diameters. Stemonitis ferruginea, Rost, (figs. 160, 161). Densely gregarious, on a well-developed hypothallus; spor- angium cylindrical, obtuse, wall thin, dark brown, evanescent; stem slender, blackish, equal to or shorter than sporangium; columella reaching to the apex of the sporangium, blackish ; capillitium dense, threads dark brown, tapering, combined to form an irregular network, the periidlieral meshes cthaut equal- ling the s2Jores in diameter; spores in the mass bright hroivn, pale brown by transmitted light, globose, smooth, 6 — 9 /x diameter. Stemonitis ferruginea, Rost., Mon., p. 196, figs. 31 — 89, 41 — 44, and 50 ; Cke., Myx. Brit., p. 47, figs. 31—39, 41—44, and 50 ; Sacc, SylL, vii., n. 1365. Exsicc. — Rav., Fung. Car., 75 (typical) ; Rav., Fung. Amer., 788. On rotten wood, &c. Britain (Lyndhurst, Highgate, Scarboro', Carlisle, Linlithgow) ; Europe ; United States ; Mexico ; S. Domingo; Cuba; Rangoon; Ceylon; Queensland; New Zealand. Usually densely gregarious, 1 — 1"5 cm. high. Readily dis- tinguished by the small peripheral meshes of the capillitium and the bright brown colour of the spores in the mass. (Rostafinski's Synonyms.) Stemonitis typhina, Willd., Ber., 408 (1787). Clathrus midus, Bolt., t. 93, f. 1 (1789). Trichia axifera. Bull, t. 447, f 1 (1791). Stemonitis fasculata, Pers,, Syn., 187 (1801). Stemonitis violacea, Schum., Saell., 1491 (1803). Stemonitis fasciculata, DC, Fl. Fr., ii., 256 (1805). Stemonitis ferruginea, Ehr., Syl. Ber., f. vi. a b (1818) ; Cooke, Hdbk., No. 1156. Stemonitis decijncns, Nees. Nov. Act. Leop., xvi. 95 (1821). Stemonitis hctcros^Jota, Oudem. Ned. Kr. Arch., i., 167 (1872). 86 A Monograph of the Mijjcogastres. Stemonitis Morgani, Peck, Sporangia crowded, stipitate, springing from a well-developed hypothallus ; cylindrical, wall evanescent ; stem much shorter than sporangium, black, shining ; mass of spores Irmon loith a fcro-^iginmcs tinge ; columella evanescent just below the apex of the sporangium ; threads of capillitium forming the network brown, suh-cgual, about 3 fA thicJc, meshes large, and about equal in size at every part, spores globose smooth, 6 — 7 \x diameter. Stemonitis Morgani, Peck, Bot. Gaz., vol. v., p. 83 ; 84tli Report State Mus., N. Y., p. 43; Sacc, Syll., vii., 1, n. 13G8. Exsicc. — Ellis and Everhart, N. Amer. Fung., Ser. II., n. 2088. On bark, wood, &c. United States. Agreeing in the large meshed capillitium with 8. Carlylci and S. maxima; differing from the former in the much smaller spores, and from the latter in the brown or sub-ferruginous colour of the spores in the mass. § § Spores loarted. Stemonitis pulchella, Bab. ' Scattered or gregarious, springing from a delicate hypothallus ; sporangic cylindrical or cylindrical-ovate, apex obtuse, base usually slightly umbilicate, wall delicate, silvery or dark ; stem rather stout, blackish, sub-equal or slightly tapering upwards, expanded at the base, shorter than sporangium ; columella reaching nearly to the apex of the sporangium ; capillitium dense, threads tapering, arcuate, brown, forming a dense, irregular net with the j7C7nj:)/ic?rtZ meshes not larger than the diameter of the spores ; spores in the mass clear ferruginous, pale brown by transmitted light, globose, minutely warted, 6 — 8 ix diameter, Stemonitis lyidchclla, Babington, Linn. Soc. Trans., 1839 ; Berk., Ann. Nat. Hist , 1841, p. 431, t, 12, f, 11 ; Cke., Hdbk., n. 1160. Comatricha pidchella, Rost., Mon., Suppl., p. 27; Cke., Brit. Myx., p. 40 ; Sacc, Syll., vii., pt. I., n. 1357. Stemonitis. 87 (Type in Herb. Berk., Kew, n. 10S41.) On twigs, herbaceous stems, ferns, &c. Britain (Bardon) ; Germany. The pcridium is sometimes ahnost white with a silvery sheen, at others brown. From 4 — 6 y. high. Much shorter and not so densely crowded as S. fnrnginca. Quite distinct from S. 2Mlcherrima, B. and C. (Rostafinski's Synonyms.) Stcmonitis 2^ulchcUa, Bab., Trans. Linn. Soc, cfr. Berk., A. and M. (1841), p. 431, t. 12, f. 11 (1839). Stcmonitis tencrrima, Curtis, Sill. Journ., 1. c, p. 849 (1848). Stemonitis tcnerrima, B. and C, Grev., n. 373 (1873). Stemonitis herbatica, Peck. Sporangia densely gregarious, stipitate, springing from a thin hypothallus, sub-cylindrical, wall evanescent ; stem usually shorter than sporangium, thin, blackish; spores in the mass hroum ; columella reaching nearly to apex of sporangium ; capillitium dense, peri2Jhcral meshes not larger than diameter of spores, threads sometimes furnished with a few short, free ends ; spores globose, very minutely tvartcd, 8 — 9 /x diameter. Stemonitis herlatiea, Peck, 26th Report State Mus., N. Y., p. 75; Sacc, Syll., vii., 1, no. 1367. (Authentic specimen from author.) On living leaves, stems, &c. United States. Often in scattered, small, densely crowded clusters, 5 — 8 mm. high. Allied to S. ferriiginea in the colour of the spore-mass, and in the small peripheral meshes of the capillitium, differing in the very minutely warted spores. Stemonitis tubulina, A. and S. Aethalium at first white, soft, 1|— 2 in. broad, 4—6 lines high, orbicular, rarely sub-oblong, basal membrane stout, silvery, pellucid, iridescent, easily removable from the substratum; surface very smooth, shining, with hemispherical warts above, 88 A Monograiih of the Myxor/astres. regularly arranged and corresponding to the apices of the component sporangia; columella (stylidia) brown, slender, capillary, aggregated, but for the most part individually free; capillitium loosely interwoven into a common mass; mass of spores brown. Stcmonitis TvJmlina, Alb. and Schu. Consp., p. 102. On decorticated pines. Germany, From the description given by Albertini and Schwoiuitz, I have very little doubt but that the organism they had in view was Siplih of the Myxogastres. Rostafinshia, Speg., Fung. Arg., Pug. III., p. 27 ; Sacc, Syll, vii., 1, p. 403 (not of Raciborski). I am not, acquainted with the single species constituting the present genus which appears to be exceptional in the septate tubes of the capillitium, &c. Distrih. Argentine Republic. Species I. Rostafinskia australis, Speg. Aethalia variable in form, thickly pulvinate — effused ; ecorticate, surface softly velvety-tomentose, at first with a longish cottony tomentum, then breaking up into powdery filaments, purple- violet or bright reddish-violet; capillitium tubes of the inter- mediate stratum cylindrical, 3 — 4 jx thick, branched, not septate, everywhere rough with minute warts, hyaline or tinged violet, tubes of inferior sterile stratum cylindrical septate, nodulosely branched, 5 — 6 [x thick, smooth, brownish ; spores globose, ovoid or irregular, filled with granular protoplasm, smooth, bright lilac, 8 — 10 x 5 — 6 /x. Rostafinskia australis, Speg., Fung. Arg., Pug. III., n. 5U; Sacc, Syll, vii., 1, n. 1381. On rotten trunks, sawdust, &c. Everywhere very common. Argentine Republic. Reticularia, Bull. Aethalia composed of numerous elongated, naked sporangia, arranged in strata and covered with a thin, common cortex ; capillitium consisting of several columella-like, slender, thin- walled tubes, that give off several lateral branches Avliich anastomose to form an irregular network. Reticularia, Bulliard, Hist. Champ., Rost., Mon., p. 240 ; Cke., Myx. Brit., p. 60; Sacc, Syll, vii., p. 418. In the present genus numerous naked, elongated sporangia, arranged in a few superposed layers or strata, are covered with an external cortex. Each sporangium has its OAvn long colu- mella, which gives off lateral branches that anastomose to form an irregular network, the branches of contiguous sporangia also Reticularia. 93 anastomose. The structure of the entire hody may be compared with the dense clusters of sporangia in Stemonitis, differing in the irregularity of the sporangia and absence of sporangial walls. Enteridium Bozeanum bears a close resemblance to the present genus, but there the capillitium is formed from the walls of the component sporangia. Several exotic species placed by Berkeley in Ecticularia belong to the genus Chromospormm. Distrih. Europe and N. America. Species 3. Reticularia lycoperdon, Host. (figs. 311, 312). Aethalium large, pulvinate, completely surrounded by a delicate cortex which varies from dull umber, through reddish- brown, to pale grey with a silvery lustre ; sometimes rough with irregular, yellowish, minute warts ; threads of the capillitium springing from the base, consisting of erect, columella-like portions with slender, anastomosing branches ; spores in the mass varying from umber to chestnut-colour, globose, about one- half the surface of the spore covered ivith a regular nctioork of liaised lines, the remainder smooth, 7 — 9 jm diameter. Beticidaria lycoperdon, Rest., Mon., p. 240, figs. 3, 4, 6, 13 ; Cke., Myx. Brit., p. 60, figs. 3, 4, 6, 13 ; Sacc, Syll, vii., n. 1424. Exsicc.—F\xc\Q\, Fung. Rhen., n. 2583. On wood, bark, &c. Britain (Bristol, Kew, Ecclesfield, Scarboro', Carlisle, Coed Coch, Linlithgow) ; Europe ; United States. Forming lumps varying from 1 — 3 inches across, more or less circular or elongated, sometimes compressed, at others pulvinate, and 1 in. or more high. Care must be taken not to confound small specimens of the present species with Lycogala e^ndendrum, which differs in the thick capillitium threads with ornamented walls. (Rostafinski's Synonyms.) .Lycogala griscnm major, Mich., t. 95, f. 1 (1729). Lycoperdon fuscuvi, Huds. Fl. Aug., 645 (1778). Mucor lycogalus, Bott., t. 183, f. 2 (1789). Beticularia lycoperdon, Bull., t. 446, f. 4, t. 470. f. 1—3 (1791). 94 A Monograph of the Mjixogastres. Lycogcda argcntea, Pers., Disp., 7 (1797). Lycogala turhinatum, Pers., Syn., 157 (1801), Strongylium fuliginoides, Ditm., t. 2, f. 1 (1809). Fidigo lycapcrdon, Schum., Saell., 1409 (1803). BcticiUaria argentca, Poir., Ency., vi., 20 (1806). Rcticularia umhrina, Fr. S. M., iii., 87 (1829) ; Corda, Ic, vi., f. 36 ; Eng. Fl., v., 308; Cooke, Hdbk., No. 1100. Sub-sect. 11. Lamprodermae. Lamproderma, Rost. Sporangia globose or broadly obovate, usually stipitate, stem continuous within the sporangium as a columella; threads of the capillitium originating from the apex only of the columella, primary branches either remaining simple for some distance or branching irregularly from the base, the branches combining to form an irregular network ; wall of sporangium usually iridescent, soon disappearing. Lamproderma, Rost., Mon., p. 202 ; Cke., Brit. Myx., p. 49 ; Sacc, Syll, vii., 1, p. 390 ; Zopf, p. 1 .56. Closely allied to Diachaca and Sicmonitis, but differing from both in having the threads of tlie capillitium originating from the apex of the columella only. Distrib. Europe; United States; Ceylon; Australia; New Zealand. Species 18. A. Spores smooth. Lamproderma violaceum, Rost. (figs. 152 — 154). Gregarious on a strongly-developed hypothallus; sporangia sub-globose, convex above, fiattmed or slightly umhilicatc Mow ; wall thin, blackish, with a dark violet lustre ; stem elongated, black, thick at the base, attenuated upwards ; columella about half the height of the sporangium, filled with large colourless cells; hrccnchcs of capillitiuni almost colourless, springing from the apex and sides of the columella, irregularly branched and com- Lawprofh'i'ma. 95 bined into a network; spores violet, globose, smooth, — 12 ju, diameter. LamiJrodcrma violacca, Rost., Mon,, p. 204, fig. G4; Cke., Brit. Myx., p. 50, f. 64 ; Sacc, Syll., vii., no. 1894. On wood, moss, &c. Britain (Shrewsbury, Castle Howard, Yorks); United States. A very distinct and beautiful species, characterized by having the sporangium flattened or umbilicate below, and the almost colourless capillitium springing from every part of the columella. From 2 — 3 mm. high. (Rostafinski's Synonym.) Stcnwnitis violacea, Fr. S. M., iii., 102 (1829) ; B, and Br., Ann. Nat. Hist., 387; Cke., Hdbk., n. 1162. Lamproderma irideum, Mass. (figs. 149 — 151). Scattered ; sporangia globose, wall thin, with steel-blue, green, or coppery metallic tints; stem elongated, blackish-brown, tapering upwards, expanded at the base into a small, circular hypothallus ; columella cylindrical, about one-third the height of the sporangium, giving origin at the apex to several thick, ascending branches, which remain simple throughout the greater 2wrtion of their length, towards the tips repeatedly bifurcating, rarely anastomosing laterally, violet-brown, not paler at the tips ; spores globose, violet-brown, smooth, 11 — 15 /x diameter. Lamproderma areyrioidcs, Var. iridea, Cke., Myx. Brit., p. 50, figs. 246—249 ; Sacc, Syll., vii., n. 1353. Mvsicc. Cke., Fung. Brit., Ser. I., 523 (as Stemonitis arcy- rioides); Cke., Fung. Brit., Ser. II., 523 (as Lamproderma areyrioidcs). (Type in Herb., Kew.) On dead leaves, moss, &c. Britain (Hampstead). Distinguished by the scattered habit, smooth spores, and in the primary branches of the capillitium remaining unbrauched except at the tips. Lamproderma suboeneum, Mass. Scattered ; sporangia globose, small, wall thin, hrovm, with a 96 A Monograph of the Myxogastres. hright co^ypery tinge, basal portion often remaining as a frill round the stem ; stem elongated, tapering upwards, expanding at the base into a minute, circular hypothallus, brown; colu- mella cylindrical, not more than one quarter the height of the sporangium, producing at the apex several primary branches, which arc at once divided into numerous arcuate hrctnchcs of equal thickness that combine to form a dense net, threads brown, often studded with organic lumps ; spores lilac-brown, globose, smooth, 1-i — 16 /x diameter. Stemonitis jjhysarioides, A. and S., Var. sulocneus, Berk., in Herb., n. 10842. On slender twigs, moss, &c. Britain (Bulmer, Yorks) ; United States. Remarkable in the capillitium consisting entirely of equal, rather stout, arcuate threads combined into a network of small, sub-equal meshes. From 2*5 — 3 mm. high, sporangium rarely attaining to '5 mm. diameter, constantly the colour of new copper. Lamproderma leucosporura, Rost. Sporangia spherical, about "5 mm. diameter, with various metallic tints ; stem black, shining, subulate, thin, passing into the sporangium as an exactly cylindrical, truncate columella; threads of the ca]_nllitium from the base variously branched and combined to form a compact nctivorh, blackish when the spores are blown away ; spores pale violet, 8 — 9 \x diameter, smooth. TjCimiyroderma leucosptprum, Rost., Mon., App., 26 ; Sacc, Syll., vii., 1, n. 1350. Lamproderma nigrcscens, Rost., Mon., p. 205 (in part). Germany. Lamproderma arcyrionema, Rost. Sporangia spherical, very small, with a silvery metallic lustre, stipitate, stem straight, subulate, black, shining, dilated into a very thin, blackish-purple hypothallus, and passing into the Sporangium as a very slender, cylindrical, truncate columella; branches of the capillitium everywhere of uniform thickness. Lamproderma. 97 divided from, the base, arcuate, variously interlaced, and combined to form a dense net withmtt free branches; spores pale violet, smooth, 6 — 7 /x diameter. Zainproderma arcyrioncma, Rest., Mon., p. 208 ; Sacc, Syll., vii., 1, n. 1352. Poland. Judging from the description, the present species appears to approach very closely such species as Stemonitis Fricsiana, B. Spores warted or cchinulate. Lamproderma echinulatum, Rost. Sporangia globose, dark steel-blue or blackish, iridescent; stem thick at the base, becoming attenuated upwards, some- times sub-cylindrical, black, filled vnth large cells that become smaller upwards; columella thick, filled with cells like the stem, about one-third the height of the sporangium, sometimes clavate; capillitium dense, originating from apex of columella and at once forming an irregular network without the usual ttndivided primary branches, tubes pale, irregular, arcuate, often flattened and triangular at the nodes; spores globose, dingy- purple, coarsely echinulate, 15 — 22 /x diameter. Lamproderma echinulatum, Rost., Mon., Append., p. 25 ; Sacc, Syll., vii., 1, n. 1344. Stemonitis echimdata, Berk., Fl. Tasm., p. 268. (Type in Herb. Berk., Kew.) Amongst moss. Tasmania. Gregarious, springing from a hypothallus, 3 — 4 mm, high. Departing from the usual type of Lamproderma in having every part of the capillitium combined to form a network with arcuate branches. Differs from Comatricha in the capillitium origin- ating from apex of thick columella only. Remarkable in having the stem and columella filled with large cells. Lamproderma Listeri, Mass. (n. sp.) (figs. 202, 203). Sporangia globose, dark-purple or blackish, iridescent, lower portion sometimes remaining as a frill round the stem ; colu- II 98 .1 Momxfraph of the Myxogastres. mella slender, about one-third the height of the sporangium, capillitium rather dense, the main branches springing from the apex of the columella which is sometimes expanded in a discoid manner, branches stout, simple for half their length, then two or three times dichotomous, axils ac^ite, secondary branches straight, sometimes anastomosing laterally, tips colourless, the other portions blackish ; stem elongated, straight or sub-flexuous, conical, smooth, blackish-brown, filled with amorphous organic matter, springing from a hypothallus ; spores globose, dingy purple, coarsely spinulose, 15 — 17 /x diameter. Scotland (Moffat) ; Tasmania ; N. Zealand. On moss, wood, &c. ; gregarious, 2 mm. high. A very fine species, remarkable for the elongated stem becoming usually much incrassated downwards, and in the apex of the columella being in some specimens expanded into a disc from the margin of which the primary branches of the capillitium originate. Both these features are characteristic of the genus Enerthenema, and the present species forms a connecting link between the last-named genus and Lamproderma. Agreeing with L. ecliinu- lata in the large, spinulose spores ; differing in the long, straight branches of the capillitium, and the stem being filled with amorphous lumps of organic matter. Lamproderma Ellisiana, Cooke. Sporangia globose, stipitate, minutely rugulose, blackish- purple, rather dull ; columella short ; capillitium originating from apex of columella, threads blackish-purple, very slender, equal throughout, repeatedly forking from the base, angles very acute; stem coloured like the sporangium and twice as long, slender above, becoming very thick downwards, and expanding into a small, circular, brown hypothallus; spores in clusters of 5 — 7 ; when the spores break up the clusters are sub-angular but soon become globose when free, pale lilac, minutely warted, 15 — 16 IX diameter. Larivproderma Ellisiana, Cke., Myx., U. States, p. 397. Badhamia p)enetralis, Cke. and Ellis, Grev., vol. v., p. 49. Lamproderma. 99 (Type in Herb., Kew.) About 1 mm. high ; scattered. " On pine boards not much decayed, white at first." (Ellis.) United States. The clustered spores probably suggested the genus Badhaniia, but the plant is a true Lam'prodermct, and quite distinct from a small species of GomatricJia, with which it is mixed. Ellis appears to have confused the true Lamprodcrma of Cooke with the ComatricJia in the N. A. F., Ser. II. Lamproderma robusta, Ellis and Everh. Sporangia globose or slightly contracted below, stipitate, blackish-purple, dull ; when in perfect condition covered with a very delicate bloom, lower portion usually remaining like a frill round the stem; columella short, thick, wrinkled; capillitium very dense, springing from apex of columella, the main branches 3 — 5 /u thick, simple for a very short distance, then h'anching and anastomosing to form a small-meshcd, very irregular dense 7ictioork ; threads tapering from the base, mostly flattened, angles often rounded, with scattered interstitial swellings, brownish-purple at the base, becoming paler upwards, attached at numerous points to sporangial wall; stem equal in length to sporangium or a little longer, blackish, smooth, equal, springing from a hypothallus; spores globose, dirty brownish-purple, minutely ivarted, 9 — 10 \j. diameter. Lamproderma robusta, Ellis and Everhart. (Described from portion of type communicated by Mr. Wingate.) On wood. United States. A very distinct and beautiful species, gregarious, springing from a common liypothallus, 2 — 2-5 mm. high. The sporangium is covered with an exceedingly thin layer of some substance resembling the bloom on a plum, which may be lime ; it cracks and breaks up into angular patches in water. Lamproderma Schimperi, Rost. Sporangia spherical, green, becoming blackish, or with a 100 A Monograph of the Mi/xogastres. reddish metallic slieen; stem black, shining, rigid, subulate, entering the peridium as an obovate columella, and about half its height; capillitium of dusky threads springing" from apex of columella, Iranches for some distance simple, then lecoming very much branched, laterally connected, and forming a dense net at the periphery, spores dingy- violet, minutely spinulose, 10 — 11 fj. diameter. Lamp)roder7iia Schim2Jeri, Rest., Mon., p. 203, fig. 63 ; Sacc, Syll., n. 1343; Cke., Myx. Brit., fig. 63. Germany. Lamproderma Sauteri, Rost. Similar to L. violaccum, but more rigid in every part ; sporangia spherical, slightly flattened below, 1 mm. broad, shining with various metallic tints ; stem black, stibulate, springing from a well-developed hypothallus; columella cylin- drical, truncate ; capillitium threads branched from the base and forming a dense network, pale when the spores are blown away ; spores dingy-violet, densely covered with sjnnules, 12 — 15 /x diameter. Lamproderma Sauteri, Rest., Mon., p. 205 ; Sacc, Syll., vii., 1, no. 1348. Tyrol. Entire specimen 2 mm. high. Lamproderma minutum, Rost. Sporangia spherical, f mm. diameter, slightly metallic ; stem black, slender, equal ; columella cylindrical, thin, truncate ; threads of capillitium colourless, rarely fasciculately branched, fascicles blending together; spores violet, delicately verruculose, 66 /x diameter. Lamproderma minutmn, Rost., Mon., Append., p. 20 ; Sacc, Syll., vii., 1, n. 1351. France. Lamproderma columbinum, Rost. Sporangia stipitate, sub-ovate or globose, blackish, reflecting Lamproderma. 101 variously coloured metallic tints ; stem slender, attenuated up- wards, longitudinally wrinkled below, purple-black ; columella cylindrical, more or less attenuated at the apex, about half the height of the sporangium ; capillitium 2mIc, 2')rimary hranclies sh(n't, hut distinct, thick, soon branching and dichotomosing in an irregular manner, combined by transverse branches to form an irregular network with numerous free, thin tips; spores globose, pale violet, minutely wartcd, 12 — 16 \x diameter. Lamproderma columhinum, Rost., Mon,, p. 203, f. 61 ; Cke., Myx. Brit., f. 61 ; Sacc, Syll, n. 1345. Laviprodcrma iridcsccns, Rost., Mon., App., p. 25 ; Sacc, Syll., n. 1346. Physarum iridcsccns, Berk., Hook. Journ., 1851, p. 20. Exsiec.—Roh., Fung. Eur., 2213; Roum., Fung. Gall. Exs., n. 1685. On moss, wood, &c. Britain (Orton Wood, Leicester; Rudloe, Twycross, Carlisle) ; Europe ; United States. From 2 — 3 mm. high, sometimes sessile and even aethalioid ; distinguished by the very short, stout, primary branches of the capillitium, the smaller branches are sometimes nodulose at intervals. (Rostafinski's Synonyms.) 3Iucor violaceus, Leess., Fl. Herb., n. 1128 (1775). Trichia violacea, Hoffm., Veg. Cr., p. 5, t. 2. f. 1 (1790). Physarum columhinum, Pers., Syst., p. 173 (1801). Trichia columhina, Poir., Encycl, no. 17 (1808). Physarum salicinum, Schum., Fl. Saell., n. 1431 (1803). Physarum hryoj^hilum, Fr., S. M., iii., p. 135 (1829). Pliysarum hryophilu7n, (3 melanocephcduvi, Cda., Ic. 1, p. 22, t. 4, f. 287 (1837). Lamproderma Saccardianum, Mass. Broadly gregarious, altogether blackish, sporangia perfectly spherical, not umbilicate, \ mm. broad, at first yellowish then opaqnc black, smooth, persistent, erect ; stem filiform, ^ mm. high, 40 ju, thick, black; hypothallus (when evident) distinct, minute, rufescent ; columella terete, half the hcis^ht of the 102 A Monograph of the Mi/.wgaHres. sporangium, apex rather obtuse ; branches of the capillitium springing from apex of columella, dichotovionsly hrancluxl, fili- form, fuliginous; spores globose, 9 — 10 /x diameter, pale smoky- violet, minutely echimdate. Lamproderma nigrcscens, Sac, Michelia, II., p. 2G2 ; Sacc, Syll., vii., 1, no. 1354 (not of Kost.). On fallen leaves of Buxus, Bobinia Binus, heaps of chips, &c, Selva, N. Italy. Distinguished from Z. 'physarioides by its smaller size and smaller spores, and from L. violaccum by the black spherical sporangium, etc. (Sacc.) Saccardo's name is antedated by Rostafinski's. Lamproderma arcyrioides, Rost. (figs. 145 — 148). Gregarious or scattered, springing from a firm hypothallus ; sporangia globose, shortly elliptical or obovate, wall thin, blackish, with blue, purple, gi'een, or reddish metallic tints ; stem usually becoming thinner upwards, blackish, shining, sometimes obsolete, passing directly into the sporangium as a short columella that breaks up at the apex into several equal, ascending branches, these again produce mtmerous lateral branches, at some distance from the base, that anastomose laterally and form a dense, irregular network, threads pale greyish- bro^vn ; spores globose, dirty violet, minutely ivartcd, 11 — 16 ju diameter. Lamproderma arcyrioides, Rost., Mon., p. 20G ; Cke., Myx. Brit., p. 50 ; Sacc, Syll, vii., n. 1353. Exsicc.—Rixh., Herb. Myc, 431; Roum., Fung. Gall, 908; Fuckel., Fung. Rhen., 1447; Rab., Fung. Eur., 797. On rotten wood, dead leaves, &c. Britain (New Forest, Scarborough, Carlisle) ; France ; Germany ; Sweden ; Denmark ; United States. From 1-5 — 3 mm. high. Near to L. violacca, but dis- tinguished by the larger and more coarsely warted spores, and in the main branches of the capillitium remaining unbranched for some distance at the base. Lamiiroderma. 103 (Rostafinski's Synonyms.) Stcmonitis arcyrioides, Somm., Tidsk. (1827); Berk., Ann. N Hist., No. 114; Cke., Hdbk., n. 1163. Stemonitis chalyhea, Pers., in litt. Stew.onitis Carestiae, Ces. and Not. (1861); Erb. Cr. Ital. 888. Stcmonitis Morthieri, Fckl., Exs., n. 1447 (1860). Lamproderma physarioides, Rost. Gregarious or scattered ; sporangia globose, wall ^m/c, with a silvery sheen, stem elongated, tapering upwards, base expanding into a small circular hypothallus, black ; columella about one*^ third the height of the sporangium, clavate ; threads of capil- litium purple-brown, springing from the clavate portion of the columella, repeatedly Ircmching in a dichotomous manner, the branches anastomosing to form a network, becoming dense and small meshed towards the periphery; spores globose, brown, minutely verruculose, 11 — 15 ju, diameter. LamprodeTma physarioides, Rost., Mon., p. 202; ficrs. 5.5, .59, 62; Cke., Myx. Brit., p. 49, figs. 55, 59, 62; Sac'c, "syll 'vii.' n. 1842. On rotten wood, moss, &c. Britain ; Germany ; U. States. Distinguished by the pale, silvery sporangial wall, the clear brown spores, and the clavate columella ; 2—3-5 mm. high. (Rostafinski's Synonym.) Stemonitis p)hysarioidcs, A. and S., Consp., t. 11, f. 8 (1805); B. and Br., Ann. Nat. Hist., no. 386; Cke., Hdbk., no. 1161.' C Sjwres rctieulated. Lamproderma Fuckelianum, Rost. Sporangia globose, almost sessile, slightly umbilieate below, reddish, metallic ; stem from its shortness almost incons2)icuous, entering the peridium for nearly half its length as a conical columella ; threads of the capillitium sparingly branched, com- bined into a loose network ; sjiores pale violet, with thin ridges combined to foim a networh, 8 — 9 /x diameter. 104 A Monopraph of the Myxogastres. Lam'prodcrma Fuckclianum, Rost., Mon., p. 207; Sacc, Syll,, vii., 1, n. 1347. Germany. Distinguished more especially by the reticulated epispore. Lamproderma Lycopodii, Raunk. Sporangia single, globose, sessile, on a violet-brown hypo- thallus. Wall, columella, capillitium, and spores violet-brown. The lower part of the wall remains, with irregular, tattered margin. Columella cylindrical, nearly reaching half the height of the sporangium. Capillitium originating singly from the upper part of the columella, in continuation forked more and more, especially at the surface of the sporangium, combined into a net by transverse branches, extremities rather colourless, Spores 12 — 18 ju, diameter, on the surface furnished ivith a delicate network of fine thickenings. Zanqirodcrma Lycqwdii, Raunk., Myx. Dan., p. 108 (in English). (Raunkier's Synonyms.) Stcriionitis cribrarioidcs, Fries, Syst, Myc, iii., p. 163; Sacc, Syll., vii., 1, no. 1370. Crihraria Lycoperdi, Nees, in litt. On the leaves of Lyeopodimn annotinum. Sealand, Enerthenema, Bowm. Sporangial wall very thin, fragile, soon disappearing; stem continuing as a columella quite through the sporangium and expanding at its apex into a discoid membrane from which the capillitium originates; capillitium of slender, sub-equal threads which bifurcate and anastomose more or less laterally, tips free ; spores free or at first in clusters. Enerthenema, Bowman, Trans., Linn. Soc. (1828), xvi,, p. 151 ; Rost., Mon., p. 209; Cooke, Myx. Brit., p. -51; Raunk., Myx. Dan., p. 02 ; Sacc, Syll, v. 7, p. 402. Enerthenema. 105 A well-marked genus, characterized by the capillitium originating solely from the peltate, flattened apex of the colu- mella; capillitium dense, threads coloured, thin, sub-equal, branching in a dichotomous or irregular manner, main branches frequently connected by transverse bars, tips free ; that is, not combined into a net, but evidently in many instances showing indications of having originally been attached to the inside of the evanescent sporangial wall. Connected with the typical structure met with in the Cohimdlifcrae by Raunkier's genus Ancyrophonis, which agrees with Encrthmema in having the columella dilated at the apex, but the capillitium originates from the columella throughout its length as well as from the disc, and the tips of the threads are not so decidedly free from anastomosing as in the present genus. Distrih. Europe ; United States ; New Granada. Sp. 3. Enerthenema elegans, Bowm. (figs. 302 — 304). Sporangia globose, with usually a minute apical umbo, wall very thin, brownish, iridescent, fugitive ; stem thick, conical, Uackish-hrrnvn, opaque, continuing as a thin columella quite through the sporangium and supporting the pendulous capil- litium from its apical disc, threads of cajnllitium hroivn, suh-equal, 2*5 — 3 /x thick, often minutely and irregularly nodulose ; spores globose, free from the first, very indistinctly verruculose, 8 — 10 /i diameter. Enerthenema elcyans. Bowman, Linn. Trans. (1828), xvi., p. 151, t. 16 ; Rost, Mon., p. 209, figs. 45, 48, 49, 52, 57. Enerthenema papillata, Rost., Mon., App., p. 28 ; Cooke, Brit. Myx., p. 51, figs. 45, 48, 49, 52, 57; Sacc, Syll., no. 1378; Raunk., Myx. Dan., p. 92, t. 5, f. 7. Britain (Wothorpe, Carlisle, Edinburgh) ; Germany ; Finland ; United States. On wood and bark. From 1-5 to 2 mm. high, wall of spor- angium disappearing very early, hence the plant is usually met with having the capillitium streaming from the discoid apex of the columella. 106 A Mo7iof/raph of the Myxogastrei^. Owing to an oversight, Encrtlienema Bcrkeleyana was intro- duced by Cooke in Myx. Brit., p. 51, as a native of Britain. Tliis species, so far as I am aware, has only been met with in S. Carolina, and is known by the clustered spores. (Rostafinski's Synonyms.) Arcyria atra, Schum., Saell, no. 1487; Fl. Dan., t. 410-i (1803). Enerthencma elegans, Bowm., Trans. Linn. Soc, xvi., p, 151, t. 16 (1828). Stemonitis mammosa, Fr., iii., 161 (1829). Stcmonitis papillata, De Bary, I.e. (1859). Encrthcnema p]iocnicolcp)ta, Bowm., msc. Enerthenema Berkeleyana, Rost. Sporangia stipitate, globose, blackish, wall evanescent ; stem black, very thick at the base, conical, becoming contracted into a thin, cylindrical, black columella that reaches to the apex of the sporangium, and there becoming expanded into a disc ; capillitium threads dark, springing from the margin of the disc, pendulous, sparingly forked, rarely joined laterally ; spores blackish-purple in the mass, at first in chisters of 4 — 14, sub- triangular at ^rst, free portion warted, 10 — 13 /x diameter. Enerthencma Bcrkeleyana, Rost., Mon., Append., p. 29 ; Cke., Myx. Brit., p. 51. (Type in Herb. Berk., Kew, no. 10888.) On boards. South Carolina. Closely resembling S. elegans, but distinct in the clustered spores that are warted on the free surface only. The stem is not always thickened below. Enerthenema muscorum, Lev. Gregarious, black ; stem setaceous, smooth, expanding at the base into a shield-like hypothallus ; sporangium smooth ; tubes of the capillitium springing from the lenticular apex of the Ancj/rophorus. 107 columella; simple for some distance, then branched in a dichotomous or vague manner; spores globose, brown. Enerthcnema musconim, Lev., Ann. Sci. Nat., Ser. IV., vol. xix., p. 289 (1863) ; Sacc, Syll., vii., 1, no. 1380. ' ' '' ' On moss. New Granada. The information respecting the present species is insufficient to indicate its specific features with exactness. Ancyrophorus, Raunk. Sporangium stipitate. Stem prolonged within the sporangium as a columella, reaching the apex of the sporangium and there expanding into a discoid membrane. Capillitium originating from this stout discoid membrane, and from the upper part of the columella proper. Threads of the capillitium only towards the extremities inconsiderably forked, the extreme branches arcuate and furnished with numerous subulate spines. Ancyro]:^]iorits, Raunkier, Myx. Dan., in Bot. Tidsskrift. (Journ. Bot. Soc. Bot. de Copenhague), 1888, p. 92, and 1889, p. 110 (in English). Clossly allied to i:7ierthenema, known by the branches of the capillitium springing from the upper portion of the stem as well as from its discoid apex. Bistrih. Denmark. Species 1. Ancyrophorus crassipes, Rauuk. Sporangia globose, stipitate. Stem shorter than the spor- angium, from an exceedingly thick base lengthened directly mto the subulate columella. Threads of capillitium combined by very few transverse branches. The extreme branches arcuate, and provided with numerous subulate spines. Stem, columella', and capillitium dull violet-black. Spores smooth or delicately warted; bright violet, 10—12 ju diameter. Ancyroiihorus emssipcs, Raunk., Myx. Dan., p. 93 (1888) and p. 110 (1890) in English. On rotten wood. Denmark. 108 A Monogmph of the Myxogastres. Raciborskia, Berl. Sporangia naked, globose, stipitate, stem elongated, entering the sporangium as a columella from one-tliird to half its height, bearing at the apex secondary, short, slender columellas, these in turn divide in a similar manner to the primary columella ; ultimate branches combined amongst themselves at every point to form a network ; extreme branchlets not free but arcuately combined with each other, Racihorslda^ Berlese, Sacc, Syll., vii., pt. I., p. 400. Bostafinskia, Racib., Myx. Crac, p. 9 (not of Spegazzini). Distrib. Poland. Species 1. Raciborskia elegans, Berl. Sporangia naked, globose, up to '5 mm. broad, stem erect, 1 — 2 mm. high, thickest at the base, then subulate-attenuated, plicate, black ; columella half to one-third shorter than the sporangium, cylindrical, 8 — 10 ju, thick; capillitium blackish- violet, exterior threads exceedingly thin, not free ; the extreme branches furnished with scattered spines; spores 9 — 10 yi diameter, obscure violet, aculeate. RacihoTsJda elegans, Racib., Myx. Crac, p. 10, fig. 5. Poland. Raciborski considers that the present species is allied to Rostafinski's genus Echiiwstdiwin. He also inclines to the idea that the genus Oiihotricha may prove to be identical with his Bacibm-slda (= Bostafinskia, Racib.).* EcHiNOSTELlUM, De Bary. Sporangia stipitate, columella absent, capillitium originating from apex of stem, tlireads arcuate, combined to form a loose network, furnished with numerous spine-like free arms. EcMnostdium, De Bary, in Rost., Mon., p. 215; Cooke, Brit. Myx., p. 53. * Hedwigia, March and April, 1889. Orthotricha. 109 Echinostelium minutum, De Bary (fig. 202). Sporangia scattered, stipitate, globose, naked, whitish; the arcuate threads of the capillitiiim springing from the apex of the stem, furnished with numerous acute, free branches; spores coloured, 6*7 — 8*3 ju, diameter, smooth. Echinostelium minutvAii, De Bary, in Rest., Mon., p. 215, figs. 53, 54, 58, and 68 ; Cke., Brit. Myx., figs. 53, 54, 58, 68 ; Sacc, Syll, vii., 1, no. 1340. Orthotricha, Wingate. Sporangium globose ; stipe elongated, entering the sporangium as a very short or obsolete columella, and then dividing into a few branches at a sharp angle. These branches fork several times, thus forming a capillitium of straight threads. The last branches meet at the surface of sporangium at a very sharp angle by twos or threes, where they are joined together by small membranaceous plates. Wall of the sporangium, with the exception of the plates and a very small collar around the stipe, not apparent. OrtJwtricha, Wingate; Proc. Acad. Nat. Sci. Phil., 1886, p. 125; Sacc, Syll., vii., pt. I, p. 400. Distrih. United States. Sp. 1. Orthotriclia microcephala, Wingate. Sporangia globose, very variable in size, from one-twelfth to one-fourth of a mm. in diameter; stipe elongated, broAvn or blackish at the base, growing lighter towards the top, more or less translucent, ten to thirty times the diameter of the sporangium in height, tapering, rugose, except at the upper part, where it more or less suddenly becomes a smooth filament, entering the sporangium as a very short, sometimes almost obsolete columella. It then divides into a few (sometimes only two) branches at a sharp angle. These branches fork several times, forming a very loose caiDillitium of straight threads, the 110 A Monograjyfi of the Myxogastres. last branches meeting at the surface of the sporangium by twos or threes at a very sharp angle, where their slightly thickened ends are joined together by minute, membranaceous plates. Sporangium wall not apparent, except a slight collar around the stipe as it enters the sporangium ; spores brown in mass, very light-violet, almost colourless, under the microscope, per- fectly smooth, 7 — 8 mm. in diameter. On rotten logs, Philadelphia, Pa. This plant has been found during three seasons in Fairmont Park, Philadelphia, Pa., in many localities. The plasmodium has a dirty-brown colour. When erecting, the dark, granular substance of the mass is left in the matter which is to form the stipe, and the globule of the sporangium becomes milky white. Before the stipe has reached its full height, say in the upper fifth, the sporangium mass leaves behind it, clustered around the stipe, several (2 — 8) clear, highly-refractive, minute globules, which, in a recently-matured plant, sparkle like dew- drops. The plant continues erecting, but from the place where the globules are left behind, the stipe very frequently suddenly narrows, sometimes to a mere filament. As the plants become old, the dew-like globules become amber-coloured, but remain transparent. These clear globules have been occasionally noted by the writer on the sporangium wall of Comatrichas, and have been considered as an indication of some degree of immaturity, hence they have not been mentioned in the description as having a specific value. In the plants, as found in different places, they are constant, though sometimes fused into one mass. The finer filaments of the capillitium, in fluid under the microscrope, are almost colourless. The plants are more or less sociable, sometimes forming patches an inch or so in diameter, and may readily be mistaken for a mould. (Wing.) Orthotricha inicrocej^hala, Wingate, in Proc. Acad. Nat. Science, Philadelphia, 1886, p. 125; Sacc, Syll., vii., pt. I., n. Ilxsicc.^EWis and Everhart, N. Amer. Fung., Ser. II., n. 2498. A very beautiful and at the same time puzzling form. I am not at all convinced in my own mind as to whether the present genus belongs to the present division or to the Fcritrichcae. Calotrirlteae. Ill CALOTRICHEAE. The varied types of ornamentation under the form of warts, spines, or raised bands, often arranged in the form of half-rings, or anastomosing to form a network on the surface of the capil- litium tubes, constitute the most pronounced character of the present order. In connection with spore dissemination, the capillitium here reaches its highest development, due to the elasticity of the tubes, which are never rigid with lime, and in the most perfect genera are quite free from the sporangial wall. The elasticity is not due to stretching, but to the sudden straightening out of previously coiled-up tubes. If an immature sporangium of Arcyria cinerca having the spores and capillitium fully differentiated, is hardened in alcohol, a section shows the tubes of the capillitium, which are combined to form a network, to be much contorted, and consequently shortened so that the net-like structure is not evident, the interspaces between the convolutions of the tubes being filled with spores. This arrange- ment of parts continues until the spores are mature, and owing to the disappearance of water, form a powdery mass, when the coiled-up tubes straighten out and the network becomes fully expanded, resulting in an increase of ten or more times in the length and breadth of the capillitium. This expansion takes place suddenly, with the result that the mass of spores are carried up and dispersed. During the expansion of the capil- litium the sporangial wall is torn into fragments, and disappears with the exception of a small portion at the base which, owing to its firmer consistency and comparative freedom from the expansion exerted by the capillitium, remains in the form of a cup or calyculus. In the genius Trichia the free tubes or elaters are very much coiled up and contorted until the spores are mature, when by suddenly straightening out, the wall of the sporangium is ruptured and some of the spores thrown out, but the arrangement is not so perfect as in Arcyria. In some of the comparatively imperfect genera, as Pcricliacna, the capillitium is scanty or obsolete. Yellow is the picdoniinating 112 A Monograph of the Myxogastres. colour of the s2:)ores as seen in the mass, passing in some species to deep orange and orange-brown ; similar colours are met with in the sjDorangial wall. In the genus Pcrichaena the inner surface of the sporangial wall is frequently covered with a layer of amorphous granules of lime. In stipitate forms of some species belonging to the genera Arcyria, Hderotrichia, and Trichia, the stem is filled with cells which are either spherical or polygonal from mutual pressure. These cells, when young, contain protoplasm, and the wall gives a cellulose reaction ; they are usually largest at the base of the stem, and become smaller upwards, passing insensibly into normal spores at the point where the stem expands into the sporangium. The tubes of the capillitium frequently pass down between these cells into the stem. It would appear as if the stem cells had originally been intended for true spores, but owing to the contraction of the base of the sporangium into a stem had been checked in their development. Arcyria. Trichia. Oligonema. I Prototrichia.- Heterotrichia. I Lachnobolus. I — Ophiotheca. Lycogala. I -Perichaena. CALOTRICHEAE. (Clathroptychium.\ PERITRICHEAE./ ANALYSIS OF THE GENERA. CALOTRICHEAE. Sub-sect. Tricheae. Elatcrs free, 7iot forming a nchoorh. Trichia. Elaters with well-developed spirals. Oligonema. Elaters without distinct spirals. Arci/riae. ii3 Sub-sect. Arcyriae. Elaters attached to wall of sporangium only slightly branched or forming a network. Prototrichia. Elaters fixed by one end to base of spor- angium, tips free. Perichaena. Sporangial wall with a layer of lime granules inside ; capillitium scanty. Ophiothcca. Threads of capillitium usually spiny, of equal thickness, forming a net with numerous blunt, free arms. Hctcrotrichia. Threads of capillitium of two distinct thick- nesses, forming a net, outer network with numerous free, pointed arms, Lachnohol'us. Threads of capillitium forming a network attached at numerous points to wall of sporangium. Arcyria, Threads of capillitium forming a network usually elastically protruded at maturity. Lycogala. Cortex containing numerous cells filled with protoplasm, capillitium tubes thick. Sub-sect. Arcyriae. Perichaena, Fries. Sporangia sessile, gregarious, dehiscing irregularly or in a circumscissile manner, wall usually double, the outer often containing granules of lime or a layer of granules of organic matter; capillitium variously developed, in some species form- ing an irregular network attached at various points to the upper portion of sporangial wall, in other species almost obsolete, threads of the capillitium without definite markings, either quite smooth or with a few scattered, rudimentary spinules or notch-like constrictions ; spores globose. Perichaena, Fries, Gast., p. 12; Rost., Mon., p. 292; Cooke, Myx. Brit., p. 77; ^opf, p. 169; Sacc, Syll, vii. pt. I., p. 420. The most prominent generic features are the circumscissile dehiscence of the sporangium, which however is not universal, the layer of amorphous particles of lime or of organic matter 114 A Monograph of the Mt/jcogastres. inside the sporaugial wall, and the smooth or obsoletely marked threads of the capillitium. The present genus is allied to Arcijria; in Arcyria paradoxa we meet with lumps of lime and of organic matter on the inner surface of the sporangial wall, but the capillitium is characteristic of Arcyria. The present genus is yet more closely allied to Oiyhiotheca ; in fact it is impossible to say where one ends and the other commences ; in Fcrichaena the capillitium threads are typically without external markings, whereas in Lachnoholus the threads have distinct raised markings, but in each genus species occur deviating from the typical idea and in the direction of the other genus, and it is doubtful whether the presence or absence of markings on capillitium threads alone, even if constant, is of sufficient value to constitute a generic distinction. Distrih. Europe ; Borneo ; W, Australia ; Canada ; United States. Species 9. Perichaena depressa, Libert, (figs. 118 — 120). Sporangia usually gregarious or much crowded, polygonal from mutual pressure, very much dcjnrsscd, and almost plane above, dehiscing in a circumscissile manner, brown, sometimes with a purple tinge, polished ; mass of capillitium and spores bright orange-ydltno ; capillitium usually well-developed, threads variable in width, smooth, rarely notched or furnished with rudimentary scattered spinules ; spores globose, minutely warted, 9 — 12 ii diameter. Perichaena depressa, Lib., PI. Cr. Ard., Fasc. IV., n. 378 ; Rost., Mon., p. 292; Cke., Myx. Brit., p. 77; Sacc, SylL, ll-S-t ; Raunk., Myx. Dan., p. 54. ^.mcc— Sacc, Myc. Ven. 500; Ellis, N. Amer. Fung., 1218; Roum. Fung. Gall, 2113; Fuckel., Fung. Rlien., 2200; Lib., PI. Crypt. Ard., Fasc. IV., n. 378. On wood, &c. Up to 1 mm. diameter. Characterized by the very much flattened sporangia and bright yellow mass of spores. Britain (Carlisle) ; Germany ; Bohemia ; Belgium ; Italy ; France ; United States. Perichaena corticalis. 115 (Rostafinski's Synonyms.) Perichaena vaporaria, Schw., Am., 2311 (1831). Ferichaena depressa, Lib., Exs., n. 378 (1837). Stcgasma deprcssitm, Cda., Ic. V., f. 13 (1842). Perichaena corticalis, Rost. (figs. 114 — 117). Sporangia spherico-depressed, crowded, dehiscing in a circum- scissile manner, lid convex, brown, bluish-purple, sometimes whitish, smooth ; mass of capillitium and spores pale ycllcno ; capillitium scanty, sometimes almost obsolete, threads thin, smooth, or here and there notched ; spores globose, warted, warts variable in size, sometimes very indistinct, at others well- developed, never smooth, 9 — 12 // diameter. Perichaena corticalis, Rost., Mon., p. 293, figs. 188 ; Cke., Myx. Brit., 78, fig. 188; Sacc, Syll., n. 1435; Raunk., Myx. Dan., p. 53. Perichaena fusco-atra, Rost., Mon., p. 294 ; Cke., Myx. Brit., 78 ; Sacc, Syll., n. 1436. Perichaena qucrcina, Fr., Gast., p. 12. (Specimen from Fries in Herb., Kew.) Exsicc. — Sydow, Myc. March., 1669; Karst., Fung. Fenn., 179; Fuckel., Fung. Rhen., 1899; Berk., Brit. Fung., 47; Desm., Cr. Fr., Ser. L, 671. On bark and wood. Britain (Lyndhurst, Kew, Derby, Castle Howard, Yorks ; Carlisle, Edinburgh, Appin, N. B.) ; Europe ; Canada ; United States ; W. Australia ; Ceylon. Very closely allied to P. depressa, if indeed really distinct as a species, the less depressed sporangium and scanty capillitium are the distinguishing features of the present species. In some specimens the wall of the sporangium has a dense layer of angular particles of lime, and is consequently nearly or quite white externally, in others the lime is scanty and in its place a layer of amorphous, coloured granules of organic matter which give the brown or reddish-purple colour to the sporangial 116 A Monograpk of the Myjcogastres. wall. The spores vary in markings from vague granulations to well-defined warts. (Rostafinski's Synonyms.) (Ferichaena coriicalis). Lycoperdon corticate, Batsch. EL, p. 155 (1783). Sphao'ocarpns sessilis, Bull., t. 417, f. 5 (1791). Trichia gymnosj^erma, Pers., Obs., vi., f. 1, 2 (179G). Tricliia circumscissa, Schrad., p. 19 (1797). Licca circuviscissa, Pers., Syn., 196 (1801). Physarum lutco-alhum, Solium., Saell., 1430 (1803). Tuhulina circumscissa, Poir., Ency., viii., 5 (1808). Pcrichacna ^^ojmlina, Fr., Gast., 12 (1817); Grev., t. 252; Eng. Fl., v., 821 ; Cooke, Hdbk., No. 1192. Perichaena pojndina, /3 sorhca, Weinm. in Fr. S. M., iii., 192 (1829). Pcrichacna qucrcina, Fries, Gast., p. 12 (1817). Trichia varia siobrufcsccns, Bong. Herb. Licca qucrcina, Wallr. Fl. Cry., 2103 (1833). Licca ccriocrcas, B. and Rav., Fungi Car., ii., 82. Pcrichacna artocreas, B. and Rav., Grev., No. 370 (1873). (Pcrichacna ftisco-atra.) Mucor lycoperdioidcs. Scop. Ann., iv., t. 1, f. 11 (1772). Trichia fusco-atra, Sibth. Fl. Ox., 1152 (1794). Sphacrocarpus sessilis. Sow., t. 258 (1803). Licea circumscissa, (3 ahictina, A. and S., p. 108 (1805). Perichaena ahictina, Fr., Gast., p. 17 (1817) ; Eng. FL, v., 321 ; Cooke, Hdbk., No. 1191. Perichaena microcar2M, Sauter. Rabh. DC. FL, No. 2180 (1844). Perichaena applanata, Mass. Sessile, adnate on a broad base, much depressed, circular in outline or aethalioid and irregular, dehiscing in nn irregularly circumscissile manner, wall with a dense layer of granules of lime on its inner surface, hright Uuc-grcy ; mass of capillitium PeyicJiaena. 117 and spores clear mrtnrjc-yclloio ; capillitiuin rather scanty, form- ing a loose net, threads 3 — 5 yi diameter, sometimes notched or with scattered rudimentary spinules; spores globose, minutely ivarted, 11 — 12 // diameter. Hcmiarcyria cvpjilanata, Cke. and Mass., Grev., vol. xvi., p, 20. (Type in Herb., Kew.) On wood. Brisbane. From 1 — 2 mm. diameter. Readily known by the bright grey sporangium and the clear orange spore-mass. The present species, owing to superficial examination, was originally described as a Hemiaoxyria. Perichaena microcarpa, Schroet. Sporangia distinct or confluent in minute clusters, generally globose, 0'5 mm.. diameter, rarely unequal, depressed, yellow- brown, smooth, dehiscing irregularly ; capillitium well-developed, threads golden-yellow, 1*5 — 2 /x thick, combined to form a loose, regular net, thickened at the angles, smooth ; spores golden-yellow, globose, 15 — 17 /x diameter, epispore strongly aculeolate. Perichaena microcarpa, Schroeter, Krypt. Fl. Schles. Pilze, p. 108; Sacc, Syll, vii., pt. I., n. 1445. On rotten leaves. Silesia. Perichaena Rostafinskii, Karst. Sporangia scattered or sub-gregarious, sessile, spherical, yellow- ish then bay, shining; capillitium ohsolete ; spores globose, smooth, or nearly so, in the mass llachish-hroum, brownish under the microscope, variable in size, 10 — 27 /x diameter. Perichaena Rostafinskii, Karst., Myc. Fenn. IV., p. 130 ; Sacc, Syll., n. 1444. On dead moss, rotten leaves, &c. Finland. The variability in the markings and size of the spores suggests that Karsten has had an immature species under consideration. Perichaena confusa, Mass. (n. sp.). Sporangia hemispherical ami scattered or aefhalioul, and. often forming an irregular nctiuorlc, pale umber or dingy ochraceous, 118 A Monograph of the Myxogastres. dehiscing irregularly; capillitium well developed, forming an irregular, loose network, threads 2 — 4 /x thick, irregularly notched; spores globose, smooth, 13 — 14 fx diameter; mass of capillitium and spores dingy ochraceous, sometimes with a suggestion of olive. (Type in Herb., Kew.) Exsicc. — Ellis, N. Amer. Fung., n. 726 (as OjMotheca ^imbrina, Berk.). On wood, leaves, &c. Britain (Lyme Regis, Yatton) ; U. States. Berkeley's description of Ojihiotheca umhrina is very imperfect, as shown by examination of his type specimen, which proves to be identical with Physarum vermiculare, Sz. Rostafinski had not seen Berkeley's type, but gave the original description and changed the name to Pcrichaena variabilis, Rost. Berkeley's imperfect description of Ophiothcca umbrina curiously agrees in many points with the present species which was unknown to him. Rarely with simple, normal sporangia, usually elon- gated and curved, forming rings, flexuous vein-like strands, or combined to form a network. The species is a connecting link between Perichacna and Ophiotheca, having the capillitium of the former without distinct markings, but the peculiar habit of Ojyhiothcca. ■ Perichaena liceoides, Rost. Sporangia globose, scattered or gregarious, dehiscing in an irregularly circumscissile manner; spores smooth, globose, 9 — 10 fx diameter ; capillitium scanty, threads slender, branched, with minute included granules. Perichaena liceoides, Rost., Mon., p. 295 ; Sacc, Syll., vol. vii., part I., n. 1438. Zicca ^;a7i7io7'«?7i, Cienk,, Pringsh. Jahrb., vol. iii., p. 407, t. 17, fig. 1 (not of Wallr.). On leaves. Germany. Douhtful species. Perichaena pallida, Berl. Sporangia gregarious, very pale tan colour; spores yellow, Lycogala. 1 1 9 but paler than in Fcrichaena australu, whose chestnut-coloured sporangia and intense golden spore-mass distinguish it from the present species. Po'ichaena pallida, Berl., Sacc, Syll., n. 1446. Stegasma pallidum, Cesati, Myc. Born,, p. 12, Borneo (Sarawak), Perichaena australis, Berl. Sporangia often circinnate, angular, central ones orbicular, at first yellow, passing through chestnut to brown, smooth, lid plane ; capillitium yellow ; spores sub-globose, granular within, opaque, yellow. Perichaena australis, Berl,, Sacc, Syll, vii,, 1, n, 1447. Stegasma australe, Cesati, Hedwigia, 1874, p, 186. Borneo. Perichaena ? pseudaecidium, Speg, Sporangia cylindrical, conical, or calyciform, 1 — 1*5 mm, long, by 0'5 — 1 mm, broad, sessile or shortly stipitate, wall very thin, cartilagineo-membranaceous, base even or minutely rugulose, dehiscing at the apex in an irregularly laciniate or fimbriate manner, chestnut or hroioiiish ; mass of spores and capillitium citrin or yellowish ; capillitium threads very slender, 1 fx thick, sparingly h-anched, here and there with solitary conical or elon- gated h^anch-lihe spines, yellowish, hyaline ; spores globose, 6 — 7 ju, diameter, often irregularly angular from mutual pressure, per- fectly smooth, pale vinous with yellow tinge, Perichaena '^.pseudaecidium, Speg,, Fung, Guar., n, 821, On living fronds of many different species of fern, and on Tillandsia muscoides. Argentine Republic. A very beautiful but paradoxical species exactly resembling a folicolous Aecidium, will probably form the type of a new genus (Speg.). Possibly a sp. of Chondrioderma. Lycogala, Mich. Sporangia aethalioid, grouped together in an intricate manner, 120 A Monograph of the Myxogastres. and forming a large plasmodiocarp, enclosed in a well-defined, thick, common cortex, enclosing near the outer side numerous cell-like masses of protoplasm ; capillitium originating at numerous points in the inner portion of the cortex, forming an irregular, loose net with numerous free ends, threads ornamented with warts or raised bands forming irregularly arranged rings, spirals, or anastomosing to form a network ; spores globose. Lycogala, Mich., Rest, Mon., p. 285; Cooke, Myx. Brit., p. 74 ; Sacc, Syll., v., 7, pt. I., p. 435. Lycoxicrdon, Bux., p. 203. Dcrmodmm, Host., Mon., p. 284; Sacc, Syll, p. 434; Cke., Myx., p. 74. The species resemble those of Reticularia in their large size, forming hemispherical or irregularly elongated masses up to four inches in diameter. The stout cortex covering the inter- woven mass of aethalioid sporangia presents a complicated structure. In L. epidendron, the most highly evolved species, the cortex contains towards the outside numerous irregularly- shaped coloured masses of protoplasm of various sizes, each surrounded by a special wall ; these are sometimes so numerous as to present the appearance of closely aggregated polygonal cells when seen in section. The tubes of the capillitium originate in the thick cortex from the cells of protoplasm just described, and frequently combine to form an irregular reticulation in the substance of the cortex ; in this position the tubes are very thick, reaching to 20 jix in diameter, and consist, as it were, of two tubes, one enclosed within the other ; the outer tube, the wall enclosing the protoplasm, has very thick, perfectly smooth, colourless walls, and disappears at the point where the capil- litium breaks through the walls to enter the mass of aethalia forming the plasmodiocarp. The inner tube has a very thin, faintly-tinged wall furnished with intermixed warts and raised bands anastomosing to form an irregular network; the inner tube is completely surrounded by the outer at its apex, until the whole structure has grown beyond the inner surface of the cortex, when the inner tube pushes through the apex of the outer smooth tube and alone forms the capillitium. The outer Lycogala. 121 common cortex cannot in the present instance be considered as excretory, and solely for protective purposes, but must be considered as a living portion of the organism giving origin to the complex capillitium, the tubes of which appear to increase in length by apical growth. The tubes of the capillitium that enter the aethalia collapse before the spores are mature, and consequently exercise no dispersive function. The scheme of classification adopted in the present work, I much regret to say, is founded mostly on characters presented by mature forms, hence the elaborate ornamention of the capillitium tubes places the present genus in the sub-family Calotriclieae ; whereas, if the capillitium is a continuation of the cortex, or rather a development directly from the cortex, Peritrichcue would be its proper position. Distrib. Cosmopolitan. Species 6. Lycogala epidendrum, Rost. (figs. 121, 122). Plasmodiocarp sub-globose, gregarious or rarely solitary, 5 — 2 cm. diameter, surface mimttely warted, variable in colour, rose- red, dingy vermilion, or brownish red ; threads of capiUitium almost colourless, 8 — 12 /u thick, springing from the cortex, becoming irregularly branched with various free, slightly in- crassated tips, now and then anastomosing laterally, furnished with warts, short bands, irregular rings or spirals, or with the raised bands anastomosing to form an irregular network ; frequently all the above modifications may be seen on the same tube, wall thin and soon collapsing ; spore-mass variable in colour, dingy purple, flesh-colour, greyish-yellow, or some- times with a dingy green tinge, spores globose, mimttely hut distinctly warted, 4 — 6 /x diameter. Lycogala e2yidendrum, Rost., Mon., 85, figs. 1, 7 — 12; Cke., Myx. Brit., 75, figs. 1, 7—12; Sacc, Syll, vol. vii., pt. I., n. 1484; Raunk., Myx. Dan., p. G2 ; Zopf, p. 1G8. Lycogala ajjine, Berk, and Br., Ceylon Fung, (type in Herb. Berk., Kew, n. 10702). Lycogala atrojmrjmreum, B. and Br., Ceylon Fung., n. 735 (type in Herb. Berk., Kew, n. 10703). 1 22 A Monograph of the Myocogastres. Lycogala nitidum, B. and Br,, Ceylon Fung., n. 734. Lycogala terrcstre, Fr., Syst. Myc, iii., p. 83 (specimen from Fries in Herb. Berk., Kew). Exsicc.—Cke., Fung. Brit, Ed. II., 614; Ellis, N. Amer. Fung., 834; Karst., Fung. Fenn., 287; Thum., Fung. Austr., 522; Fckl., Fung. Rhen., 1475; Rav., Fung. Car., 78; Mong. and Nest., n. 85; Kunze, Fung. Sel., 197; Westendorp, Crypt. Belg., 741 ; Jack, Leiner u. Sitz., 380 ; Desmaz. Cr. Fr., Ser. I., 609 ; Rab., Fung. Eur. 2140 ; Sydow, Myc. March., 186 ; Roum., Fung. Gall., 2813. On decaying trunks, stumps, &c. Plasmodium rose-coloured, or sometimes bright scarlet. Britain (Kew, New Forest, Rudloe, Scarboro', Carlisle, Appin, N. B., Abergavenny); Europe; U. States ; Bermuda ; Venezuela ; Cuba ; Algeria ; S. Africa ; Himalayas; Ceylon; Madagascar; New Guinea; W. Australia. (Rostafinski's Synonyms.) Fungus coccineus, Ray Syn., ii., 386 (1690). Fungus sangtdneus, Bocc. Mus., p. 304 (1697). Fungus non vcsms, Lses. Fl. Prus., 96 (1703). Lycoperdon sanguineum J' ^.u^iip. Fl. Jen., 304 (1718). Bovista miniata, Dill. Cat. PL, 197 (1719). Lycoperdon e'pidcndron, Bux, Hall, p. 203 (1721) ; Fl. Dan., t. 720; Light, f. Fl. Scot., No. 1068; Holms. Ot., t. 31; Bolt, t 119, f. 1; Sow., t 52; Purt Midi. FL, p. 701; RehL Cant, p. 566 : With. Arr., iv., p. 457. Lycogala glohosum^ Mich., t. 95, f. 2 (1729). Lycoperdon sanguineum, Buxb., t. 29, f. 2 (1740). Lycoperdon sphaericum, Gled. Meth., 150 (1753). Mucor iii. sphaericus, Gled. Meth., 161 (1753). 3fucor secundus, Schff., t 193 (1762). Lycogala sessile, Retz., Ac. Holms., 254 (1769). Mucor lycogala, Scop. Cam., ii., 1645 (1772). Mucor fragiformis, Schff. Bar., No. 283 (1774). Lycoperdon variolosum, Huds. FL Ang., 645 (1778). Lycoperdon epijihyllum, Huds. FL Ang., 645 (1778). Lycogala. ] 23 Lyco]perdon pysiformc, Jacq. Misc., t. 7 (1788). Galepcrdon epidendron, Wigg. Fl. Hols?it. 1148 (1780). Lycoperdon chalyhettm, Batsch. El., p. 155 (1781). Zycoperdon verrucosum, Batsch, El, p. 155 (1781). Reticularia rosea, DC. Bullet. Phil, f. 8 A, B, c (1798). Lycogala miniata, Pers. Obs., ii., 26 (1 790) ; Grev. S. C. Fl, t. 38 ; Nees., t. 8, f. 97 ; Gray. Arr., i., 568. Lycogala punctata, Pers. Syn., p. 158 (1801). Lycogala plumhca, Schum. Saell, 1408 (1803). Lycogala fcrniginca, Schum. Saell, 1406 (1803). Reticularia miniata, Poir.,'Ency., viii., 22 (1808). Reticularia punctata, Poir., Ency., viii.. No. 21 (1808). Reticularia rosea, Poir., Ency., viii.. No. 4. Lycogala miniata, Johnst. Fl Berw. Lycogala cpide7idribm, Fr., S. M., iil, 80; Berk. Engl Fl, V. 307; Cooke, Hdbk., No. 1095. Lycogala ajffinis, B. and Br., Ceylon Fungi, No. 735. Lycogala conicum, Pers. Plasmodiocarp sessile on a broad base, often attenuated towards the apex, hence more or less conical, shining, purple, or pale ochraceous-brown, at maturity minutely wartcd or scurfy ; mass of spores ochraceous-olive, sometimes with a tinge of flesh- colour; tubes of capillitium numerous, originating from the inner portion of the cortex, 8 — 10 ju diameter, almost colourless, frequently branched and anastomosing laterally, free ends numerous, markings variable, warts or irregular raised bands more or less anastomosing or sometimes almost obsolete ; spores globose, minutely wartcd, 4 — 6 /x diameter, Lermodium conicum, Rost., Mon., p. 284; Sacc, Syll, vil, pt. I., n. 1483. Gregarious, '5 — 1 cm. or more in diameter. From a careful examination of specimens of the present species that have been examined and named by Rostafinski, I am satisfied that the genus Lermodium is synonymous with Lycogala. In both cases the leading idea is a vein-like plas- modiocarp covered with a common cortex, the outside portion 124 A Mono(jf?Yfph of the Myxogastres. containing numerous cell-like aggregations of protoplasm, while the inner portion gives origin to the tubes of the capiUitium which originate from the cell-like masses of protoplasm. Rostafinski's generic diagnosis of JDcrmodium is altogether misleading, inasmuch as it implies that we are dealing with a single sporangium, whereas in Rostafinski's own specimens the plasmodiocarp structure is distinct. Furthermore it is very doubtful as to whether the present species is anything more than Lycogala epidcndrum. It is certain that Lycogala atropur- picoxum, B, and Br., and Lycogala nitidum, B. and Br., referred to the present species by Rostafinski, both belong to Lycogala epideiidrum. On rotten wood. Sweden ; Germany ; Ceylon. (Rostafinski's Synonyms.) Lycogala conica, Pers., Syn., 159 (1801). Lycogala conicum^ Fr., S. M., iii., 82 (1829). Lycogala atroj^uriyurettm^ B. and Br., Ceylon Fungi, II., n. 735. Lycogalc nitidum, B. and Br., Ceylon Fungi, II., n. 734. Lycogala flavo-fuscum, Rest. (fig. 123). Plasmodiocarp hemispherical or elongated, 3 — 9 cm. diameter, cortex smooth or slightly wrinkled when dry, yellow-brown, bright chestnut-brown, or greyish-umber ; mass of spores greyish- umber or with a flesh-coloured tinge; tubes of capiUitium rather scanty, almost colourless, 4 — 5 /x tliich, slightly rugidosc 01- with indistinctly raised lands, sparingly branched ; spores globose, very minutely wartcd, 5 /x diameter. Lycogala jiavo-fuscum, Rost., Mon., p. 288; Cooke, Brit. Myx., p. 76; Sacc, SylI.,.no. 1485 ; Zopf, p. 167. On stumps and trunks. Britain (King's Cliffe); Europe; United States ; Ceylon. A large species, usually solitary, distinguished from L. epi- dcndrum by the smooth cortex and the scanty capiUitium with thinner tubes and rudimentary ornamentation. Li/cogala. 125 (Rostafinski's Synonyms.) Di^jhtherium flavo-fiiscurii, Ehr., Syl., p. 27 (1818). Eetimlaria fiavo-fusca, Fr., S. M., iii., p. 88 (1829) ; B. and Br., Ann. Nat. Hist., No. 591. Edicularia testacca, Wallr., Fl. Germ., 209 (1833). Lycogala ochraceum, Mass. (n. sp.). Plasmodiocarp sessile, sub-globose, gregarious, springing from a firm, hroadly ejpused kypothallus, ochraceous, at first smooth, becoming minutely furfuraceous ; mass of spores very pale ochraceous; capillitium scanty, tubes sparingly branched, with a few indistinctly liaised bands, 5 ix diameter, collapsing ; spores globose, 3' 5 — 4 \x diameter, minutely and sjmrsely vcrrueulosc. On rotten wood. Baliia ; Java. (Challenger exped.) (Type in Herb. Berk., Kew.) Often densely gregarious and variously angular from mutual pressure, at other times larger and scattered ; from '5 — 2*5 cm. diameter. Lycogala minutum, Sacc. and Paol. Plasmodiocarp gregarious, superficial, sessile, base narrow, globoso-depressed, 4 — 5 mm. h^oad, smooth (not punctate), at length with crowded, very minute depressions, yellow, becoming ochraceous ; hypothallus scanty, white, mucedinous ; tubes of capillitium filiform, short, sulsimplc, hyaline ; spores globose, aspcrulose, 3 ix diameter, yellow, then colourless. Lycogala minutum, Saccardo and Paoletti, ' Mycetes Mala- censes' Atti del R. Instit. Veneto di scienze, lettere, ed arti. Tome VI., Ser. VI., p. 5, t. 5, f. 1 ; Sacc, Syll., n. 1486. On fallen decorticated rotten branches. Malacca. Apparently closely allied to Lycogala ochraceum, but distin- guished by the scanty mucedinous hypothallus and in the cortex not becoming furfuraceous. Lycogala rufo-cinnamomeum, Mass. (n. sp.). Plasmodiocarp hemispherical, gregarious, "5 — 1 cm. diameter, surface smooth, ochraceous or greyish, some times with a tinge 126 A Monogrcqyh of the Myxogastres. of red ; mass of spores bright reddish-cinnamon ; threads of capillitium colourless, much branched, vjolls smooth, not collapsing when dry, 4 — 5 \x thick ; spores globose, 10 ju diameter, rather coarsely warted. On fragments of rotten wood lying on the ground. South Africa. (Type in Herb. Berk., n. 10706.) Gregarious. Distinguished by the dense capillitium, the tubes of which have thick, smooth walls, and the large spores. Prototrichia, Rost. (emended). Sporangia simple or fasciculate, stipitate or sessile, thin, reflecting metallic tints, dehiscing irregularly ; capillitium well- developed, elaters attached by one end to the basal portion of the sporangium, becoming attenuated upwards and dividing into a variable number of slender, tapering, free tips, the slender branchlets are sometimes more or less connected laterally, elaters furnished with spirals, irregular rings, or entirely smooth. Prototrichia, Rost., Mon. App., p. 38; Cooke, Myx. Brifc., p. 65; Sacc, Syll., vii., pt. I., p. 437; Mass., Revis. Trich., p. 349. Trichia, B. and Br., Ann. Nat. Hist., Ser. III., vol. xviii., p. 56, pi. 2., f. 4. Coo^nuvia, Rost., Mon., p. 289 (in part); Cooke, Myx. Brit., p. 76 (in part) ; Sacc, Syll., vii., pt. 1, p. 289 (in part). Alwisia, B. and Br., Joum. Linn. Soc, xiv., p. 87, t. 2, f. 6, and XV., t. 2, f. 1 ; Mass., Mon. Trich., p. 348. Trichia, Rost., Mon., p. 246 (in part) ; Sacc, Syll, vii., pt. I., n. 1494 (in part). As above defined the main features of the genus are the thin iridescent sporangial wall and the elaters 'attached at one end to the base of the sporangium, and becoming divided above into a variable number of tapering, free tips, the branchlets are rarely connected transversely, but never to such an extent as to suggest a network. Distrib. Europe ; Tasmania ; Ceylon : 5 species. Prototrkhia. 127 A. Spores smooili. Prototrichia flagellifera, Rost. (figs. 130—132). Sporangia scattered, globose, attenuated below to a narrow point of attachment, wall thin, smooth, copper-colour, and reflecting metallic tints ; mass of capillitium and spores flesh- colour; elaters 7 — 9 ix broad at the base, tapering towards the apex, branching at some distance towards the point of attach- ment into two or three arms, each generally again divided near the apex, spirals thin, not prominent, sometimes crowded, at others distant, disappearing below the ultimate branchlets, brown, becoming colourless towards the tips; spores globose, smooth, 10 — 18 IX diameter. Prototrichia flag ellifer, Rost., Mon. Appendix, p. 38 (in part); Cooke, Myx. Brit., p. 65 ; Sacc, Syll., n. 1492 (in part) ; Mass., Rev. Trich., p. 850, pi. vii., f. 23. Trichia fiagellifer, B. and Br., Ann. Nat. Hist,, Ser. III., vol. xviii., p. 56, pi 2, f. 4 (no. 1143). Dermatricha fiagellifer, Cooke, MS. (Type specimen in Herb. Berk., Kew, n. 10,905), On dead wood. Britain (Badminton, Gloucester, Carlisle) ; Sweden. Scattered, or rarely 3 — 5 in a cluster, but not crowded together, '5 or a little more in diameter. A very neat species, the sporangial wall is unusually persistent, hence the plant is generally found in good condition. The smooth spores and elaters thickest at the point of attachment mark the species. Prototrichia metallica, Mass. Sporangia scattered, stipitate or sessile on a broad base, spherical or depressed and lenticular, smooth, shining, copper- colour with metallic tints ; stem very short, rather thick, darker in colour than the sporangium ; mass of capillitium and spores pale fiesh-colour or yellowish ; elaters elongato-fusiform, 6 — 7 [i at thickest part, 300 — 400 /x long, tcrminatiiig at the apex in a 128 A Monograjyh of the Myxogastres. peticil of simiile or branched, cylindrical, smooth, somctiuics nodu- lose filaments, 2 \i thick, and 40 — 60 ju, long; spirals broad, flat, close ; spores globose, smooth, 9 — 1 1 /x diameter. Prototrichia metallica, Mass., Rev. Trich., p. 350, fig. 26. Trichia metallica, B. and Br., Fl. Tasm., p. 268. Prototrichia fiagcUifera, Rost., Mon. Append., p. 38 (in part) ; Sacc, Syll., n. 1492 (in part). Prototrichia clcgantula, Rost., Mon. Append., p. 39, fig. 246 ; Sacc, Syll, n. 1491. (Type in Herb. Berk., Kew, n. 10,905 a.) On wood, Tasmania; Sweden, Sporangia "5 — 1 mm. diameter. Rostafinski founded his Prototrichia clcgantula on a specimen in the Berkeley Herbarium at Kew, which was sent by Fries, and marked " {Pcricliacna ?) nova species, in Betula, Lindblad." This specimen on examin- ation proves to be identical with Trichia metallica, B. The last-named species is given as a synonym of P. fi.agcllifcr, B. and Br., by Rostafinski, the outcome of superficial examination. Prototrichia bombarda, Mass. Sporangia broadly fusiform or elliptical, several seated on the apex of an elongated stem, smooth, darh-hrown, sometimes with a purple tinge ; stem same colour, hollow, springing from a well-developed hypothallus ; mass of elaters and spores brown ; elaters cylindrical, 6 — 7 /a thick, sometimes furnished with one or two swollen portions, walls thin, collapsing when dry, un- hranchcd, with a few scattered, very rudimentary spinules, and a very indistinct open spiral, free ends obtuse, rarely attenuated and bifid ; spores globose, smooth, 5 — 6 ^x diameter. Ahoisia ho^nharda, B. and Br., Journ. Linn. Soc, xiv., p. 87, t. 2, f. 6, and xv., t. 2, f. 1. Trichia fragilis, Rost., Mon., p. 246 (in part) ; Cke., Brit. Myx., p. 63 (in part) ; Sacc, Syll, n. 1494 (in part). On decayed wood. Gongolla Forest ; Ceylon. (Type in Herb. Berk., Kew, n. 10921). From 2 — 3 mm. high, scattered. The fasciculate sporangia PrototvicJiia. 120 are in some specimens connate excejot at the tips, and then present the appearance of a single sporangium with several subacute apical lobes. The stem is somewhat contracted and wrinkled longitudinally when dry. The present plant resembles to a certain extent, when examined with a pocket-lens, fasci- culate forms of TricUa fragilis, and from such superficial examination Rostafinski gave it as a synonym of the last- mentioned species. If it had been properly examined this mistake would not have been made. B. Spores warted. Prototrichia cuprea, Mass. (figs. 127 — 129). Sporangia scattered or rather crowded, sub-globose, usually sessile on a broad base, rarely attenuated below, or with a very short distinct stem, bright copper-colour, shining, sometimes iridescent; mass of capillitium and spores reddish flesh-colour; capillitium copious, elatcrs G— 8 /x thich at ■point of origin from wall of sporangium, 60—80 ^ long, then branching once or twice in a dichotomous manner, branches tapering upwards, 150—200 [I long, each ending in a corymhosc tuft of slender, smooth, colourless filaments, 1—2 /x tliiek, main trunk and branches brownish, with rather close, not prominent spirals; spores globose, minutely warted, 10—13 jix diameter. Prototrichia cupora, Mass., Rev. Trich., p. 351, pi. vii., f. 24. (Type in Herb., Kew.) On dead thorn. Scarborough ; Carlisle. Sporangia pure white when immature, -5—1 mm. diameter, bright copper-colour, polished and often iridescent, especially when old and empty. Characterized by the warted spores and the slender filaments terminating the main branches of the elaters, which are not always so densely corymbose as figured in Rev. Trich., pi. vii., f. 24. A form of the present species sent by Mr. Camm from Smethwick has the elaters without a trace of spiral marking, thus approaching 1\ chamacleontina. 130 A Monoriraph of the Myrogastres. Prototrichia chamaeleontina, Mass. Sporangia spherical or spherico-depressed, attached by a narrow base, wall very thin, highly iridescent, dehiscing irregu- larly; capillitium well-developed, threads thick at the point of attachment, tapering upwards, often nodulose, without external markinc/s, breaking up into several free, pointed spines that are sometimes connected hy to'cmsvcrsc hands, dingy brown ; spores pale pink, globose, minutely wartcd, 9 — 11 ^x diameter. Cornuvia metallica, Kost., Mon. App., p. 35 ; Cooke, Myx. Brit., p. 76 ; Sacc, Syll, n. Fhysarum metcdlicum, Berk., IVtag. Zool. and Bot., No. 29, t. 3, f. 8 ; Cooke, Myx. Brit., p. 16. (Type in Herb. Berk., Kew.) On twigs, bark, dead leaves, &c. Britain (Clifton, Notts ; Carlisle, Smethwick). Scattered or in groups of 2 — 3, about 1 mm. diameter. Sporangial wall very iridescent, reflecting green or rosy tints, mass of spores pinkish or flesh- colour. The previous existence of a species of Frototrichia called mctcdlica, accounts for the change of name in the present species. Ophiotheca, Curr. (emended). Sporangia sessile with single or double wall, frequently con- taining amorphous particles of lime, either regular, circular, or depressed, with circumscissile dehiscence, or aethalioid, or form- ing a vein-like, anastomosing plasmodiocarp ; capillitium form- ing a loose network, attached to the wall at various points, and usually with free arms, threads with spines, warts, or short bands irregularly scattered ; spores globose. Ophiotheca, Currey, Quart. Journ. Micr. Science, vol. ii., p. 240. Pcrichaena, Rost., Mon., p. 292; Cke., Brit. Myx., p. 77; Sacc, Syll., vii., 1, p. 420 (all in part). Cm^nuvict, Rost., Mon., p. 289 ; Cke., Brit. Myx., p. 76 ; Sacc, Syll., vii., 1, p. 423 (in part). Opiilotltcca. 131 Closely allied on the one hand to Perichacna, and on the other to Arcyria. Differs from the latter in the sessile, much compressed sporangia often becoming aethalioid, presence of lime in the wall, and the capillitium with numerous free arms and attached to the wall at many points. From Pcricliciena in the threads of the capillitium being in some way ornamented. Lachnobolus differs in the stipitate sporangia. Didrih. Europe ; United States ; Cuba ; Ceylon. Species 9. A. Threads of rcjdllitmm spinnlosc. Ophiotheca circumscissa, Curr. Sporangia either regular, circular, depressed, or dehiscing in an irregularly circumscissile manner, or becoming aethalioid, sinuous, and often combined into a network ; wall smooth, chestnut, or brown ; mass of capillitium and spores yclloic ; threads of caiyiUitium 2 — 4 }i tJiicIc, with scattered, straight, or hent spines, 1 — 2 ju long, combined to form an irregular network, with numerous free arms often slightly incrassated at the tips ; spores globose, smooth, 8 — 19 /x diameter. Olihiotheca chrysospcrma, Curr. Quart. Micr. Journ., v., ii , p. 240, t. 9, f. 1—5. Cornuvia circumscissa, Rost., Mon., p. 290 ; Cke., Myx. Brit., p. 76; Sacc, Syll., n. 1452; Schroeter, Kr. Fl. Schles., 108. Cornuvia dictyocarpa, Crupa, Cosmos, Lemberg, 18C6; Sacc, Syll., n. 1453. On bark. Britain ; Germany ; France ; Switzerland. (Rostafinski's Synonyms.) Lignidium quereinuni, Fr., Stirp. Femsp., 83 (1825). Trichia circumscissa, Wallr., Fl. Cr. Germ., n. 2219 (1833). Arcyria glomcrata, Fr., Sun. Veg. Scand., 457 (1849). Ophiotheca chrysosp)crma, Microsc. Journ., p. 240, t. 9, f. 1 — o (1854), Trirhia Curreyi, Crouan, 1. c, 16 (1867). 132 A MonograpJi of the Myjcogastres. Far. scahra, Schroeter, Kr. FI. Schles. (Pilze), p. 1 09. Threads of capillitiuni with rigid spines, 1 — 2 /x long. Silesia. Var. sjnnosa, Schroeter, Kr. Fl. Schles. (Pilze), p. 109. Threads of capillitiuni with scattered acute spines, 5 — 6 jj. long. Silesia. Ophiotheca Wrightii, Berk, and Rav. (fig. 197). Sporangia circular, much depressed, dehiscing in a circum- scissile manner, or becoming aethalioid and irregularly vein- like; wall bright or blackish -brown, smooth, with granules of lime inside ; mass of capillitium and spores Iright, deep yellow, sometimes with a flesh-coloured tinge; capillitium well de- veloped, forming a loose, irregular network, the threads are densely and very minutely vemoculose, and in addition are furnished with numerous straight or ficxuous sharp-pointed spines, 2 — 4 /x long ; spores globose, minutely wartcd, 10 — 14 u diameter. OjJhiothcca Wrightii, Berk, and Curt., Linn. Journ., vol. x., p. 349. Gornuvia Wrightii, Rost., Mon. App., p. 36 ; Sacc, Syll., vii., 1, n. 1454. Exsicc. — Fung. Cub. Wrightiani, 540. (Type in Herb. Berk., Kew, n. 10888.) On wood. Cuba ; Ceylon. Very closely allied to 0. circumscissa, more especially the Var. spinosa, but distinguished by the warted spores. OpMotheca irregularis, Mass. Sporangia spherical, much depressed, rather convex above, usually gregarious, dehiscing in an irregularly circumscissile manner, wall bright-brown, containing lime inside; mass of capillitium and spores bright yellow ; capillitium well-developed, forming a loose net, threads 3 — 4 /x diameter, equally covered with distinct slender spines 'b — 1 ju lung ; spores globose, minutely verriiculose, 9 — 11 /x diameter. Ophlotheca. 133 Pcrichacim irrcf/ularis, Berk, and Curt., Grev., vol. ii., p. G8 ; Sacc, Syll., n. 1439. (Type in Herb. Berk., Kew.) On bark. Lower Carolina. Sporangia up to 1 mm. diameter, often much crowded and consequently irregular in outline. Resembling Pcricliaena deiJressa in general appearance, but distinguished by the distinct spines uniformly and rather thickly scattered over the capillitium threads. Ophiotheca nitens, Mass. Sporangia single or few gathered in clusters, gIoho^c.-2niriform , sessile, or with a short stem, splitting irregularly, greyish-brown with a fine violet tone of metallic lustre, '5 mm. broad. Wall single, almost without particles. Tubes of the capillitium long, slightly branched, by irregular enlargements fastened to the wall, unequally and very delicately sinnulose, equally broad throughout, 1 — 1*5 /x broad. Spores delicately spinidose, yellowish, hyaline, 10 — 12 jx diameter. Pcrichaena nitens, Raunkier, Myx. Dan., p. 107, t. 2, f 11, On bark of oak and fir, and on decayed leaves of pine. Denmark (Sealand, Jutland). B. Threads of elaters with raised lands which may he short and wart-like, or comhined to form rings or a nctworJc. OpMotheca reticulata, Mass. (figs. 199, 200). Plasmodiocarp venulose, sinuous or anastomosing to form an irregular network, dingy ochraceous-umber, wall containing numerous masses of lime inside ; mass of capillitium and spores pale ochraceous-umber; capillitium scanty, threads forming a very loose network, 3 — 5 /x thick, densely covered with short, straiglit, or curved raised hands and rounded, warts ; spores globose, luith minute, scattered urtrts, 13 — 14 fj. diameter. Licea reticulata, B. and Br. Pcrichaena reticulata, Rost., Mon. App., p. 35 ; Sacc, Syll., vii., 1, n. 1442. 134 A Monograph of the Myxogastres. (Type in Heik. Berk., Kew, n. 10852.) Ou bark. Ceylon. Resembling in many respects Ophiothcca vcDiiicuIaris, but distiuguislied by the very minute, scattered warts on the spores. OpMotheca cano-flavescens, Mass. Sporangia gregarious, standing on a thin, yellow-grey hypo- thallus, globose to hemispherical, or lengthened to rcniform, sessile, '5 mm. broad, bright yellow-grey, splitting more or less regularly with a lid. The wall contains numei'ous round or angular particles. The upper part of the wall, on its inner side, provided with delicate thickenings combined into a regular network ; meshes of the network 5 — 6-gonal and 12 fx diameter. Civpillitium scarcely evident, only consisting of a few, slightly- branched or quite unbranched threads of variable width, un- cc[ually warted, colourless, yellowish ; without regard to the thickenings the tubes are 1"5 — 2 ju broad. Spores golden-yellow, delicately warted, 12'5 — 14 /x diameter. Perichacnct cano-Jlavesccns, Raunkier, Myx. Dan., p. 107, t. 2, fig. 10. On bark of beech. Denmark (Lolland). OpMotheca vermicularis, Mass. (figs. 198, 198 «). Sporangia circular, irregularly lobed, or venulose and anasto- mosing to form an irregular network ; when the sporangia are circular or simply lobed there is usually a well-defined, up- raised marginal portion, wall smooth with masses of lime insitle, varying from flesh-colour, through brown to umber; mass of capillitium and spores flesh-colour or bright-yellow ; capillitium well-developed, attached at numerous points to wall of spor- angium, combined to form a loose, irregular net, threads 3— 4 ix diameter, cqv.ally covered vjith mimerous sliort, straight or curved, sometimes branched, raised hands ; sometimes a few rounded warts are intermixed, these latter now and again run out with prominent spines; spores globose, densely and thichly covered njiih small, dejrrcssed vxtrts, 10 — 13 /x diameter. O pit lot her a. 135 Physarum vermicnlarc, Sz., Syn. Fung. Amer., n. 229G. (Specimen from Schweinitz in Herb. Berk., Kew.) Perichaena vermicularis, Rost., Mon, App., p. 84; Sacc, Syll., vii., 1, n. 1437. Ophioiheca umhrina, B, and C, Grev., vol. ii., p. 68 (1873). (Type in Herb. Berk., Kew.) (Not Perichaena variahilis, Rost., Mon., p. 205 ; Sacc, Syll, n. 1440.) Perichaena artocreas, B. and Rav., Grev., v. ii., p. 68 (1873). (Type in Herb, Berk., Kew). Perichaena corticalis, Rost., Mon., p. 293 (in part) ; Sacc, Syll, n. 1435 (in part) ; Cke., Myx. Brit., p. 73 (in part). Perichaena Friesiana, Rost., Mon., 296 ; Sacc, Syll., n. 1441. On rotten wood and bark. United States, In Schweinitz's specimen there is a clear transition shown from normal perithecia to the venulose, reticulate plasmodiocarp. In the non-venulose form the margin is uniformly raised all round. The threads covered with short raised bands, looking like warts under a low power, and the spores densely covered with depressed warts are very characteristic, Ophiotheca anomala, Mass. Sporangia scattered or gregarious, sessile, sub-sphaeroid, dingy- ochraceous, rather shining, about 1*5 mm. diameter ; threads of the capillitium 4 — 6 /x thick, cylindrical, ends numerous, truncate, often terminated hy clavately swollen tips, icith crowded raised hands arranged as circles; spores globose, smooth, dingy-ochra- ceous, under tlie microscope very pale tinge of yellow, 6 — 7 /x diameter. Cornuvia anomala, Karst., Myc Fenn., 4, p. 131 ; Sacc, Syll., vii., 1, 1455. Trichia anomala, Karst., Not. Siillsk. pro Faun, et Flor. Fenn., ix. (1868), p. 354. Ophiotheca serpula, Mass. Plasmodium venulose, yellow; mass of spores and ca])initium golden-yellow; capillitium threads cylindrical, 3'3 ju Ihid; ■iriih 136 A Monograph of the Myxogastres. raised hands in the fcn-ia of o-ings, '4^ — "7 /it high, combined into a net with free, more or less swollen tips; spores globose, 10 /it thick, with raised hands forming a j^ohjgonal nctworl-. Arcyria serpnla, Wigand, Pringsh. Jahrb., vol. iii,, p. 44, t. 3, f. 18. Cornuvia serpula, Rost., Mon., p. 289 ; Sacc, Syll., n. 1451. On wood. Germany. Wigand describes the spores as areolate on the surface, but judging from his figure of a spore, mag. 720 diameters, it appears that the areolation is due to raised bands combined to form a polygonal network. OpMotheca pallida, B. and Curt. Sporangia Jlcxuous, 'paU ; threads of capillitium yellow, nodu- lose, minutely echinnlate ; spores globose, "0004 inch in diameter ( = about 11 ii). 02?hiotheca 'pallida, Berk, and Curt., Journ. Linn. Soc, vol. x., p. 350 (1869). On dead stems of herbaceous plants. Upper Carolina. The flocci are very irregular in outline, and have far shorter processes than the other two species. [6*. chrijsospcrma and C. Wrightii] (M. J. B.) There is no specimen in the Berkeley Herbarium. Lachnobolus, Fries. Sporangia stipitate, sessile, or aethalioid, dehiscing irregularly or by the disappearance of the upper evanescent portion of the wall ; capillitium combined into a net, which is attached at numerous points to the wall of the sporangium ; spores globose. Lachnoholns, Fries, Flor. Scand., p. 356 ; * Sacc, Syll, vii., p. 433; Cooke, Brit. Myx., p. 74; Zopf, p. 170. Very closely allied to Arcyria; in fact the only important point of difference consists in the present genus having the capillitium attached by arms at numerous points to the wall of the capillitium, hence it is not protruded elastically when Lachnuholus. 137 mature as an aid in spore dispersion, a function performed by the elastic capillitium in some species oi Arcyria, but not all, as in several species belonging to the last-named genus, the capillitium is in many places attached to the wall, as in A. chrysosiwra, A. imllidula, &c. In Arcyria the markings on the capillitium threads are generally more developed than in Lachnoholus. Distnh. Europe ; United States. Species 4. Lachnobolus globosus, Rest. (fig. 204). Sporangia globose, stipitate, lower half of qjorangial wall stout, persistent, upper half very thin, eva^iesccnt, yellow, becoming whitish; stem about equal in length to sporangium, straight, coloured like the sporangium, filled with large irregular angular cells; mass of capillitium and spores colour of ground ginger; capillitium dense, combined into an irregular network, attached at many points to persistent portion of sporangium, threads varying from 3 — 6 ^ broad, closely covered with minute warts ; capillitium not elastic ; spores at first sub-angular from mutual pressure, then becoming globose, but the thick wall remains slightly thickened at the angles, hence looking as if furnished with a few rudimentary warts, 6 — 7 ju. diameter. Lachnoholus glohosics. Rest., Mon., p, 283 ; Sacc, Syll., vii., pt. L, n. 1479. On dry involucres of sweet chestnut. United States. (Specimen from Schweinitz in Herb. Berk., Kew.) Gregarious, about 15 mm. high. (Rostafinski's Synonyms.) Arcyria glohosa, Sz., Consp. Fung. Carol., n. 400 ; A.mcr. Fung., n. 2340 (1822). Cratcrium globosum, Fr., S. M., iii., 154 (182D). Nassula glohosa, Fr., Summ. Veg. Sc, 4o() (1849). Var. minor, Ellis, N. Amer. Fung. Exs., n. 1397. 138 A Mono9) • Hall pi 1, f. 6 (1742). ' ■ \ - J' 'V Clathrus 2Jcdiculatus, Guett., Obs. 1, 16 (1747). Arcyria pedimla, Hill, p. 47 (1751). Clathrus demidatus, Linn., Sp. PI., 1179 (1753). Tridiia, Hall, No. 2164, t. 48, f. 6 (1768). Mucor clathroidcs, Scop. Carr., ii., 492 (1772). Mucor injriformis, Leers. Fl. Herb., 1135 (1775). Clathrus pcdunculcikis, Batsch. FL, p. 141 (1783). Lyco2icrdon rufttm, Dicks Cry., fasc. i., p. 25 (1785). Stcmonitis dcnudata, Rehl. Fl. Cant. (1786). Emlolus crocatus, Batsch, t. 30, f. 176 (1786). Stcmonitis crocata, Willd. Fl. Ber., 1189 (1787). Stcmonitis coccinea, Roth. Fl. Germ., i., 548 (1788). Triclna dcnudata, Vill. Fl. Dauph., 1060 (1789) '• Purt Mid Fl, t. 24,f. 2; Sow., t. 49. Trichia graniformis, Hoffm. V. Cr., i., p. 3 (1490). Trichia cinnabaris, Bull, t. 502, fig. 1, b. c. (1791). Stcmonitis crocca, Gmel Sys., 1467°(1791). Trichia rufa. With. Arr., iii., p. 478 (1795). 144 A Monograph of the Myxoijaatve^. Arcyria punicca, Pers. Disp., p. 10 (1797); Eng. Fl., v., 318; Cooke, Hdbk., No. 1170 ; Fung. Brit., ii., 520. TricJda 2yu7jmrca, Schura. Saell., 1472 (1803). Arcyria rufa, Sebum. Saell., 148G (1803). Arcyria melanocqiliala, Solium. Saell., 1484 (1803). Arcyria conjugata, Sebum. Saell., 1485 (1803), Arcyria cincta, Sebum. Saell., 1480 (1803). Arcyria cylinclrica, Sebum. Saell., 1486 (1803). Trichia cinnaharis, DC. Fl. Fr., No. 688 (1805). Arcyria f'lisca, Fr. Gast., p. 17 (1818). Arcyria vcrnicosa, Rost., Mon. Supp., p. 36. Arcyria punicea. Var. cribroides, Raunk. Tlie inner mesbes equal and of ordinary widtb ; on tbe surfaee of tbe capillitium, on tbe contrary, groups of very small and single laj^ers sueeeed eaeb otber alternately. Raunk., Myx. Dan., p. 108, t. 3, f. 12. Jutland. Arcyria ferruginea, Rost. (figs. 250 — 253). Sporangia globose or ovate, sometimes deformed, stipitate, dull vermilion or rusty-red, sometimes pale reddisb-pink ; stem tbin, length variable, usually short, filled witb large cells ; mass of capillitium and spores brick-red, brownish, sometimes oebra- ceous ; capillitium becoming elastically protruded, threads com- bined to form a dense, irregular network, viorc or less acutely triangttlar, one side with imrallel iirojccting ridges, the two others vnth a fine, irregular network ; spores globose, smooth, 9 — 1 1 fx diameter. Arcyria ferruginea, Rost., Mon., p. 280, fig. 194; Cke., Myx. Brit., p. 73, fig. 194 ; Sacc., Syll, vii., 1, n. 1470. Arcyria intricata, Rost., Mon. Suppl, p. 72. Exsicc. — Fuckel, Fung. Rben., n. 1444. On rotten wood. Britain (Hampstead, Carlisle, Scarboro') ; Europe; United States ; Ceylon. Distinguisbed amongst tbe red species of Arcyria by tbe Arcyria. 145 portion of the capillitium threads not occupied by tlie pro- minent ridges being ornamented with a dehcate network of slightly raised lines. (Rostafinski's Synonyms.) Arcyria fcrruginea, Sauter., p. 316 (1811). Arcyria latcritia, DeBy. Mycet., 24 (1859). Arcyria fcrruginea, Fckl. Sym., p. 337 (1869); Berk, and Br., Ann. Nat. Hist., No. 1603 ; Grevillea, v., p. 13. Arcyria incarnata, Rost. Sporangia crowded, ovate or sub-globose, stem very short or absent, wall reddish, often with a brown or yellow tinge ; mass of spores and capillitium flesh-colour, rosy, or reddish-brown, sometimes ochraceous ; ccqnllitium attached to the apex of stem- wall, and ivith very few attachments to hasal portion of thin, sporangial wall, threads combined to form a dense, elastically protruding network, with many free, fattened, or swollen sjnnose tips, markings in the form of half-rings or spines, or a mixture of the two arranged in a loose spiral, and mixed with small, spinose warts, there are frequent swellings in the length of the threads; spores globose, smooth, pinkish or tinged with ochre, 6 — 8 ju, diameter. Arcyria incarnata, Rost., Mon., p. 275, figs. 187, 199 ; Cooke, Myx. Brit., p. 71, figs. 187, 199. Arcyria adnatcc, Rost., Mon. Supp., p. 72. M'sicc. — Fuckel, Fung. Rhen., 1442. On rotten wood and bark. Britain (Lyndhurst, Kew, King's Cliffe, Norths, Carlisle, Aboyne, N. B.) ; Europe ; United States ; India ; Ceylon. Usually crowded, sub-sessile. To — 2 mm. high. The wall of the sporangium is thinner, and the permanent base little or not at all longitudinally ribbed ; the capillitium threads are generally round, but sometimes compressed. Closely allied to A. jmnirra, of which the present is perhaps only a variety; intermediate stages between the two ai'e not wanting. 146 A Monorfraj>h of the Afijjcoijastres. (Rostafinski's Syuonyms.) Clathroidcs pmjmTeurn, Mich., t. 94, f. 2 (1729). Arcyria scssilis, Hill, p. 47 (1751). Clathrus ramosus, a., Retz. Vet. Hand,, 253 (1769). Lycoperdon hypoxylon, Pallas., ii., 503 (1771). Clathrus adnatus, Batsch., p. 141 (1783). Stemonitis trichia, Roth. Fl. Germ., 1,, 549 (1788), Stemonitis lilacina, Schr,, Fl. Bav., ii., 1784 (1789). Stemonitis incarnata, Pers., in Gmel., 1467 (1791). Trichia cinnabaris. Bull, t. 502, f. 1, D (1791). Arcyria incarnata, Pers., Obs., t. 5, f. 4, 5 (1796) ; Eng. Fl., v., 318; Cke., Hdbk., 1171. Stemonitis carnea, Trent., 222 (1797). Stemonitis glohosa, Trent., p. 222 (1797). Trichia fiexuosa, Schum. Saell., 1465 (1803). Arcyria lilacina, Schum. Saell,, 1476 (1803). Arcyria carnea, Schum, Saell, 1477 (1803). Arcyria dcntata, Schum, Saell., 1478 (1803), Arcyria rosea, Spr. Arcyria incarnata, ^fiexuosa, Fr., S. M., iii., 179 (1829). Arcyria nutans, /3 cxigua, Bong., in Weinm., 609 (1836). Arcyria fiextiosa, Rabh., Fl. Cry., 2158 (1844). Arcyria irregularis, Racib. Sporangia pinkish-red, stipitate ; stem up to '5 mm. high, filled with large, colourless cells; calyculus hemispherical, mcm- hrane mi7iutely ivarted ; capillitium forming an irregular net, meshes of variable size; threads flattened, margin undulate, from 4'5 — 9*1 ju wide, 2-5 — 4*5 thick, threads everywhere densely covered with irregular obtuse cones 0*8 — 1*5 /x broad, for the most part elongated; spores 7 — 7*5 jix diameter, membrane firm, smooth, almost colourless. Arcyria irregularis, Racib., Myx, Crac, p. 15, fig. 10; Sacc, Syll., vii., 1, n. 1406. Poland. Ai-rf/rid. 147 Arcyria (Erstedtii, Rost. Sporangia ovate, in the typical form, stipitate, stem sliortcr than sporangium, firm, erect ; like the sporangium, dirty brown ; mass of capillitium and spores hrown, often ivith a red tinge; capillitiura forming a rather dense net, threads cylindrical, 3 — 5 /x diameter, loitli scattered, elliptical, swollen portions, spar- ingly and equally covered with slender spines of various lengths ; spores smooth, 8 — 10 /x diameter. Arcyria (Erstedtii, Eost., Mon., p. 278, fig. 19G; Sacc, Syll., vii., 1, n. 1468 ; Raunk., Myx. Dan., p. 60, t. 8, f. 1 ; Cke., Myx. Brit., fig. 196. On wood. Britain (Burnham Beeches, Bucks) ; Denmark ; Russia. Characterized by the net, without free arms, having its threads swollen at intervals, and sparingly and equally covered with slender spines of variable length, the longest about equal to diameter of thread. Arcyria umbrina, Schum. Sporangia single or gregarious, on a thin, shining hypothallus, ovate, stipitate. Receptacle on the inner side provided with a network of delicate thickenings. Stem brownisli-black, longer than the sporangium. After dehiscence of the sporangium the capillitium enlarges itself but slightly. Capillitium and the mass of spores ochre-brown. Tubes of net of the capillitium with thickenings in the form of half-rings encircling the thread in a spiral, without regard to the thickenings 2'5 — 3 // broad. Spores smooth, 5 — 7 /u, diameter. Arcyria umhrina, Schum., t. 3, f. 5. Scliumacher : Fl. Saell., No. 1479; Fl. Dan. Tab., 1975, fig. 1; Raunk., Myx. Dan., p. 108, t, 3, f. 8. On rotten stumps. Summer ; autumn. Near Copenhagen. When comparing the two Arcyria — species delineated in the Flora Danica, 1975, figs. 1 and 2, with the descriptions of them, it will be seen that the text to fig. 1 in reality belongs 148 A Monogra])h of the Myxogastres. to fig. 2, which no doubt is Arcyria cincrca, what also may be seen by comparison of this figure with the authentic figure in Bulliard's Hist, des champ., &c., Tab., 447, fig. 3. This error is to be found in all authors who have cited Tabula 1975 of Fl. Dan. I beg to observe that I follow the figure no. I. of the Tabula and correct the text according to this, con- sequently for A. cinerea I have cited Fl. Dan. Tab. 1795, fig. 2. Fig. 1 on the Tabula — fig. 2 in the text — Arcyria umhrina, Schum., is hitherto quoted as synonymous with A. pomiformis, Roth., but I think they are rather different both in their shape and in their colour. On a rotten stump near Copenhagen I have found the above described Aoryria, which throughout harmonizes with the figure named and with Schumacher's description of his A. iimlrina. This species I therefore believe to have again found. As may be seen from the description, A. umhrina is in the thickenings on the tubes of the cajDillitium very different from A. pomi- formis, and more connected with A. inmicca. (Raunkier, in Myx. Dan.) From the above it appears that Raunkier considers he has collected the species intended by Schumacher. It Avill be noted, however, that no type specimens are factors in the determination, but only figures. Arcyria insignis, Kalchb. and Cke. Sporangia scattered, very minute, not more than -5 mm. high, cylindric-ovate, rosy or vermilion, upper portion falling away, calyculus minute ; stem half as long as sporangium, similarly coloured, filled with large, globose, coloured cells which pass into normal spores at the apex; capillitium dense, protruded elastically, threads cylindrical, thick-walled, combined into an irregular network ; everyiuhere densely and equally covered loith stout, acute spines about 1 [x high ; spines not connected into a network at the base; spores globose, smooth, 11 — 12 /x diameter. Arcyria insignis, Kalchbr. and Cke., Grev. x., p. 143; Sacc, Syll., no. 1475. On rotten wood. S, Africa. Arcyria. 149 (Type in Herb., Kew.) Distinguished amongst the red species by the cylindrical threads of the capillitium being equally and densely covered with short spine-like warts or very short bands. Arcyria fusca, Fries. Every part unibo^-brown, sometimes with a tinge of olive ; sporangia stipitate, globose or very hroadly elliptical ; basal por- tion remaining as a calyculus ; stem 3 — 4 times as long as sporangium, very thin, equal, filled with large cells that become smaller upwards ; capillitium protruded elastically, attached to base of sporangium, the threads passing down between the cells of the upper portion of the stem, threads round, combined to form a dense network, with a single row of warts or very short 2'>lates arranged in a very open spiral; spores globose, smooth, 7 — 9 /x diameter. Arcyria fusca, Fries, Gast., p. 17. Arcyria punicea, Rost., Mon., p. 268; Cke., Myx. Brit,, p. 69; Sacc, Syll., vii., 1, no. 1457 (all in part). Exsicc. — Fuckel, Fung. Rhen., 1443. (Specimen from Fries in Herb. Berk.) Pn wood. Britain (Batheaston) ; Sweden ; Germany. Gregarious, 3 — 4 mm. high. Considered by Rostafinski as synonymous with Arcyria punicea, from which it differs in the ornamentation of the threads, consisting of spines or short flattened plates not occupying more than \ diameter of thread, long, thin stem, colour, &c. Arcyria versicolor, Phill. Sporangia elongato-pyriform or clavatc, often attenuated downwards into a short, thin, weak stem ; wall of sporangium smooth, polished, varying from bright clear yellow to dark olive ; dehiscing irregularly, basal portion persistent; springing gre- gariously from a well-developed hypothallus ; mass of capillitium and spores clear yellow to dusky olive ; capillitium attached only to inner surface of stem-like base of sporangium; forming 150 A Morwgrcrph of the Myxogastres. a dense, irregular network, protruded elastically ; threads cqiudly covered in every part %vith short, variously curved, and frequently anastomosing, slightly raised hinds; sometimes a more or less perfect ring occurs; spores globose, smooth, 8 — 10 /x diameter. Arcyria versicolor, Phillips, in herb. Arcyria versicolor, Phill., Grev., v., p. 115, t. 88, f. 8, a — e; Sacc, Syll., vii., 1, n. 1473. Arcyria vitellina, Phill,, Grev., v., p. llo, t. 88, f. 7, a — e; Sacc, Syll, n. 1474. On tiunks. California. A beautiful species, 3 — 3'5 mm. high. Variable in colour, but constant in all important characters. I am indebted to Mr. Phillips for an oj^portuuity of examining the type specimens. Arcyria nutans, Rost. (figs. 140, 141). Sporangia cylindrical, sometimes ovate, very shoiily stipitate, wall dingy yellow ; stem same colour, filled with large cells ; capillitium and mass of spores pale opaque yellow; eapillitium elastically protruded, elongated, drooping, threads combined to form a dense, irregular network, thickenings in the form of prominent half-rings, often with projecting spines arranged .in a diffuse spiral, the remainder covered loith a fine, irregular net- woQ'h ; spores globose, pale yellow, smooth, 7 — 8 fx diameter. Arcyria nutans, Rost., Mon., p. 277 ; Cke., Myx. Brit., p. 72. Exsice. — Fuckel, Fung. Rhen., n, 1439. On rotten wood and bark. Britain (Kew, Scarboro', Carlisle) ; Europe ; United States. Scattered or gregarious, 1'5 — 2 mm. high. Somewhat re- sembling A. cinerea, but distinguished by the short stem, the elastically protruded, very long, drooping capillitium, which is usually the colour of ground ginger, but sometimes tinged with brown, and more especially by the portion of the capillitiam threads not occupied by the half-rings being ornamented with a fine network of slightly raised lines. A irifvia. ] 5 1 (Rostafinski's Synonyms.) Clathroidcs fiavescens, Mich., p. 214 (1729). Clathroides longissimum, Hall, p. 10 (1742). Clathrus ramosus, 13. Retz. Vet. Handl., 253 (17G9). Trichia nutans, Bull, t. 502, f. 3 (1798) ; Sow., t. 260. Stemonitis nutans, Gmel., Sys., 1467 (1791). Arcyria flava, Pers. Obs., i., 85 (1796) ; Grev., t. 809. Stemonitcs amaena, Trent., p. 222 (1797). Trichia clongata, Schuni. Saell., 1464 (1803). Arcyria ahdacca, Sclmm. Saell., 1474 (1803). Arcyria nutans, Grev. Fl. Ed., p. 455 (1824); Eng. Fl., v., 319 ; Cooke, Hdbk., No. 1173. Arcyria ^aliens, Wallr. in litt. Arcyria straminea, Wallr. Crypt. Germ., 2232 (1833). Arcyo'ia nutans, c, minor, Bong., in Weinm, p. 609 (1836). Arcyria cinera, Mass. (figs. 137—139, and 248, 249). Sporangia varying from globose to cylindrical, stipitate, simple or digitato-fasciculate on a common stem, wall thin, smooth, grey, or yellowish, cup-like base persistent after de- hiscence ; stem slender, aboiU equal in length to sporanginiji or longer, straight, colour of sporangium, filled with large cells that become smaller upwards; mass of capillitium and spores yellowish-grey or grey ; capillitium dense, [irutriiding elastically, and remaining erect, threads combined to form an irregular network, variable in thickness, often smooth or minutely warted towards the base, the remainder equally covered with close-set short spines, or half-rings or spines arranged in a diffuse spiral ; spores globose, smooth, rarely minutely granular, 7 — 9 /x diameter. Arcyria cinerea, Rost., Mon., p. 272, figs. 182 — 185, 193; Cooke, Myx. Brit., p. 71, figs. 182—185, 193; Sacc, Syll., vii., 1, n. 1459. Arcyria 'j-jomiformis, Rost., Mon., p. 271 ; Cke., Myx. Brit., p. 70. Arcyria stricta, Rost., Mon. Sujjp., p. 71. 152 A Monogiutph of the Mihrogastres. Arcyria Friesii, B. and Br., Ann. Nat. Hist., n. 1G02 (type in Herb, Berk., Kew) ; Cke., Myx. Brit., p. 71, figs. 251 and 259. Exsicc. — Fuckel, Fung. Khen., 1440; Sydow, Myc, March., n. 1499; Rab., Fung. Eur., 369 ; Ellis, N. Amer. Fung., 1115. On wood and bark. A variable species, from 1*5 — 4 mm. high, the capillitium remains erect after expansion. In the fasciculate form the sporangia are cylindrical and elongated, 2 — 8 springing from a long, slender stem. Britain (Lyndhurst, Swansea, Kew, Buhner, Scarboro', Car- lisle, Appin, N.B.); Europe; United States; Brazil; Venezuela; S. Africa ; Himalayas ; Ceylon. (Rostafinski's Synonyms.) (Under A. cinerea.) Clathrus recutitus, Linn. Sp., 1649 (1764). Trichia, Hall., No. 2166 (1768). Trichia cinerea, Bull, t. 477, £ 3 (1791). Stcmonitis recutita, Gmel. Sys., 1467 (1791). Stemonitis cinerea, Gmel. Sys., 1467 (1791). Arcyria albida, Pers. Disp., t. 1, f. 2 (1797). Stemonitis glauca, Trent., p. 221 (1797). Arcyria cinerea, Schum. Saell., 1480 (1803) ; Fl. Dan., t. 1975, f. 1 ; Eng. FL, v., 318; Cooke, Hdbk., No. 1172. Trichia carnea. Wall, in litt. Arcyria camea. Wall. Fl. Germ., 2234 (1833). Arcyria trichioides, Corda Ic, ii., f. 86 (1838). Stemonitis grizca, Opiz. in Lotos, 215 (1855). Arcyria iKillida, B. and C. Grev. (1873), No. 365. Arcyria stricta, Rost., Mou. Supp., p. 36. (Under A. pomiformis.) Stemonitis X)oriiiformis, Roth. Fl. Germ., i., 548 (1788). Stemonitis ochroleuca, Trent, in Roth., 221 (1797). Stemonitis lutea, Trent., 221 (1797). Arcyria umbrina, Schum. Saell., 1479 (1803); Berk. Ann. N. Hist, No. 389; Cooke, Hdbk., No. 1174. Arcyria. ' 153 Arcyria silacea, Ditm., t, 8 (1817). Arcyria ochroleuca, Fr. S. M., iii., 181 (1829); Berk. Ann. N. Hist., No. 115; Cooke, Hdbk., No. 1175. Arcyria hitea, Scliwz. Fung. Car., 2339 (1831). Arcyria globosa, Weinm, teste Fr. Stemonilis ochracca, Opiz. herb. Arcyria cinerea. Var. cribroides, Raunk. Tubes of the capillitium broader than those of the type species, the inner meshes equal and of ordinary width, on the surface of the capillitium, on the contrary, groups of very small meshes and single larger succeed each other alternately. Raunk., Myx. Dan., p. 107. On decayed branches of fir. Denmark. Arcyria digitata, Rost. Scattered or gregarious, several s^wrangia, S—7 , fasciculaic on a common stem; sporangia shortly cylindrical, apex obtuse, basal cup-like portion persistent; mass of capillitium and spores pale opaque greyish-yellow, or with an ochraceous tinge; stem elongated, equal, rather slender, expanded at the base into a cn^cular hypothallus; capilHtium dense, hecoming very much elongated after dehiscence, threads forming a network, meshes very irregular in size ; here and there minutely warted, varying m thickness ; spores globose, sinooth, 7—10 )u diameter. Arcyria digitata, Rost., Mon., p. 274; Sacc, Syll vii 1 n. 1460. Stemonitis digitata, Sz., Syn. Amer., n. 2350. (Specimen from Schweinitz in Hb. Berk.) Arcyria Zeprieurii, Mont., Ann. Sci. Nat., Ser. IV., vol. iii., p. 141. (Type in Herb. Mus., Paris.) Arcyria hicolor, B. and C, Journ. Linn. Soc, vol. x.,'p. 349. (Type in Herb. Berk.) Exsicc.~Vm,g. Cub. Wrightian., 542 (as Arcyria hicoLr, B. and C). On woud and bark. Cuba; U. States; Cayenne. 154 A Monogrrtph of the Mi/xogastres. From 3 — 4 mm. high ; several sporangia fasciculate on a common stem, rarely simple. Closely allied to Arcyria cioierea, differing in the larger size of the sporangia. Sometimes the capillitium and spores are brownish or red. (Rostafinski's Synonyms.) Stcmonitis digitata, Sz., Amer., f., n. 2350 (1834). Arcyria Lepricurii, Mont., Ann. Sci. Nat., v. iii., p. 141, no. 603 (1855). Arcyria Cookei, Mass. (n. sp.). Gregarious ; every part hluish-grcy ; sporangia narroirly cylindrical, elongated, wall very thin except the basal portion which persists as a shallow calycnlus ; stem erect, elongated, very slender, eo^a], filled toith irreynlar lumps of organic matter ; capillitium dense, becoming protruded elastically during de- hiscence and remaining erect, innermost threads very thin, 1*5 — 2 // smooth, combined to form a very large-raeshed, irregular network, outer threads 4 — 5 ix thick, densely and uniformly covered with minute warts, forming a small-meshed network ; spores globose, smooth, 8 — 9 /u diameter. On wood. Brazil. (Type in Herb., Kew.) From 3"5 — 4 mm. high ; stem about equal in length to sporangium. Placed by Dr. Cooke along with Arcyria cinerea, from which it differs in the thin, elongated sporangium, the details of the capillitium, and in the thin long stem being filled with amorphous lumps of organic matter. Arcyria dictyonema, Rest, Sporangia ovate, stipitate ; stems sjJO'inging in clusters from the substratum, capillitium not very much divided, threads cylindrical, width variable, generally 3'5 /n thick, siih-equally covered loith an elevated net^cork, the hands of which are sjnnulose, spines 1 — 2 ix high ; spores smooth, 9 — 10 /x diameter. Arcyria. 155 Arcyria didyonema, Rost., Mon., p. 279, fig. 195; Cke., Myx. Brit., fig. 195; Sacc, SylL, vii., 1, n. 14G9. Russia; Germany. Appears to approach very closely some forms of A. cmcrea and A. nutans. (Rostafiuski's Synonyms.) Arcyria nutans, (a) sordide ochracea, Bong., in Wein., 1. c, p. 609 (1836). Arcyria ochracea, D. By., MS. Arcyria cinnaviomca, Kaulfuss, MS. Arcyria Hariotii, Mass. (n. sp.). Gregarious or much crowded and irregularly polygonal from mutual pressure, depressed, sessile on a broad lase, sporangial wall primrose-yrlloiv, smooth, shining, capillitium and mass of spores dingy yellow ; threads of capillitium 6 — 8 ix diameter, forming a dense, irregular net, uniformly and densely covered with mimUe warts ; spores globose, wall thick, synooth, 7 — 9 fx diameter. France. Communicated by M. Hariot of the Pai-is Museum. On decayed wood. About "5 mm. diameter, superficially closely resembling Oligoncma nitcns. Agrees in the smooth spores, dense capillitium, and polished sporangium with Arcyria versicolor, but in the latter the sporangium is clavate or elongato- pyriform, and the markings of the capillitium consist of short irregularly arranged bands. Arcyria aurantiaca, Raunk. Sporangia gregarious, ovate, or shortly cylindrical, stipitate ; stem same length or shorter than sporangium. Thickenings on the inner side of the receptacle, in the form of fine warts ; wall, capillitium, and mass of spores brick-red to orange. Tubes of the capillitium with irregularly connected, close-standing, ring-like thickenings, 5 — 7 /x broad; spores smooth, 10 — 11 /x diameter. lo6 A Monograjih of the Mi/jcogastres. Arcyria aurantiaca, Rauuk., Myx. Dan., p. 109, t. 3, figs. 4, and 9—11. On rotten wood, Denmark. From the above description it is difficult to indicate either the prominent features of the species intended, or its affinity with other species. * * Spores luartcd. Arcyria similis, Racib. Sporangia shortly stipitate, cherry-coloured ; threads of capil- litium thick-walled, 4*2 — 5 — 8 yi. broad, cylindrical or a little compressed, everywhere ivith plate-elevations; plates generally in the form of half-rings not arranged in a regular spiral manner, hut irregularly, or overlapping, or in spirals irregularly broken up; spores 8.3 /u. diameter, almost colourless, wall thin, dis- tinctly verruc^Uose. Arcyria similis, Racib., Myx. Crac, p. 13, f. 8; Sacc, Syll., n. 1462. Poland. Arcyria Raciborsckii, Berlese. Sporangia stipitate, stem filled with cells; calyculus hemi- spherical, membrane thin, reddish, densely covered on the inner surface with very short, acuminate warts ; capillitium at the nodes and often also on the internodes inflated into globose vesicles, forming a loose net ; threads of lower portion of the capillitium flattened, one margin with incrassated teeth, remainder of threa/is smooth, the remainder and greater portion having the threads sich-cylindrical, with the teeth arranged in a spiral, the remainder of the thread with undulating ridges, in a varying number of spirals, here and there becoming indistinct, or anastomosing to form an irregular network on the surface of the thread ; spores vcrrucidose, 10*5 — 11*6 /a diameter. Arcyria BacihorscJcii, Berlese, Sacc, Syll., vii., 1, n. 1467. Arryria. 157 Arcyria decvpiens, Racib., Myx. Crac, p. 16, f. D (not of Berkeley). Poland. Arcyria inermis, Racib. Sporangia stipitate, brick-red ; stem filled with cells of various sizes, the interspaces filled with capillitium threads ; capillitium forming a network, threads cylindrical or a little flattened, 4*2 — 10*8 ix hrosid, protuherances comhincd into a nettvorh, meshes 1"6 — 2 u long and almost equally broad ; spores 9"9 — 10'8 \x. diameter, membrane firm, reddish, distinctly vemiculose. Arcyria inermis, Racib., Myx. Crac, p. 14, f. 7 ; Sacc, Syll., vii., 1, n. 1465. Poland. Position uncertain, o^uing to absence of information ixspecting the spores. Arcyria bonariensis, Speg. Sporangia minute, | — f mm. long, J — \ mm. broad, in densely crowded clusters of 5 — 20 individuals, bright citrin-ycUow ; stem half as long as peridium and same colour ; tubes of the capillitium springing from the hollow of the stem, equal, 3 ju, thick, densely micricate, very pale yellowish-green ; spores globose, protoplasm granular or guttulate, wall rather thick, 10 // diameter. Arcyria bonariensis, Speg., Fung. Arg., Pug. III., n. 90 ; Sacc, Syll., vii., 1, n. 1471. On old worked wood. Argentine Republic Allied, to A. mitans, but distinct. (Speg.) Arcyria affinis, Rost. Sporangia cylindric-ovate, 1 mm. high, stem of about equal length to sporangium, gregarious; mass of capillitium and sjjores purple-ferruginous, dull orange-red ; threads of capillitium forming a network, lower meshes o'6, upper 5"4 fx broad, mark- ino-s as in C. incarnaia. 158 A Monograph of the Myxogastres. Arcyria affmis, Rost., Mon., p. 276 ; Sacc, Syll., vii., 1, n. 1463. On trunks. Sweden ; Germany. (Rostafinski's Synonyms.) Arcyria vcrmicularis, Schum., Fl. Saell., n. l-iTo, teste Fr. (1803). Arcyria pimicca, ^ vcrmicularis, Fr., 1. c, iii., p. 178 (1829). Arcyria cinnamomea, Hazsl. Ginnajnon-colour inside and out; sporangia gregarious, at length distinct, cylindrical ; stem equal to sporangium in length, diaphonous-white ; tubes of capillitium combined into a hexa- gonal network, 3 — 4 //, thich, with sliwt, cylindrical, diaphonaus warts; spores cinnamon. Arcyria cinnamomea, Ha,z&\., Just., Bot. Jahr., 1877, p. 156; Sacc, Syll., vii., 1, 1477. Arcyria decipiens, Berk, Gregarious, sporangia stipitate, obovate, ochraceous, capil- litium free above, obovate, erect, with the mass of spores ochraceous ; spores globose. Arcyria decipiens. Berk., Fung, of Darwin's Voy,, Ann. Nat. Hist,, v., 4, p. 447, t. 10, f. 3, On palm stem. Rio Janeiro. Habit of Trichia clavata, but capillitium reticulated, and thread without spirals (Berk.), No specimen exists in Berkeley's herbarium, B. Hcmiarcyria. Thrcaxls of capillitium ivith continuous ridges spirally arranged. § Spores minutely warted. Arcyria rubiformis, Mass. (figs. 262 — 264). Sporangia olovate-ollong, usually fasciculate on a common ridged or contorted stem of variable length, sometimes sub-globose Arcf/ri'f. 159 and sessile on a hypotliallus, rarely an irregular plasmodiocarp, hrown or almost black, polislied and with a metallic lustre or opaque ; mass of capillitium and spores orangc-lroicn ; threads of capillitium orange-brown, 8 — 9 /u thick, combined into a dense elastic net, which at maturity elongates considerably, carrying the apical portion of the sporangium at its apex, where it remains in the form of a cap, free tips numerous, usually terminated by from one to three short, smooth spines, rarely obtuse, spirals rather distant, furnished with numeroiis slender sphies ; spores globose, very minutely verruculose, 10 — 12 /x diameter. a. genuina; sporangia cylindrico-turbinate, dark-brown, opaque, or with a steel lustre, seated on a common fasciculate stem. ;8. sessilis ; sporangia sessile, cylindrical or sub-angular from mutual pressure. y. plasmodiocarpa ; irregular, attached by a broad base to a hypothallus. Hemiarcyria ruUformis, Rost., Moo., p. 262; figs. 201, 230, 231; Cooke, Myx. Brit., figs. 201, 230, 231 ; Schroeter, p. 114; Sacc, Syll., vii., n. 1512; Raunk., Myx. Dan., p. 63, t. 3, f. 15, t. 4, f. 6 ; Mass., Rev. Trich., p. 353, fig. 31 (in part). Exsicc.—Roxun., Fung. Gall, 1686 ; Fckl, Fung. Rhen., 1438 (as Trichia ruhiformis) \ Cooke, Fung. Brit., 612 (as Trichia Neesiana) ; Jack, Leiner u. Sitz. Krypt. Bad., 421 (as Trichia ruhiformis) ; Sacc, Myc. Ven., 962 (as Trichia ^jyj-^/brwiis) ; Hemiarcyria Ellisii, Mass., Rev. Trich., p. 354, f. 30; Karst., Fung. Fenn., 700. A widespread and very beautiful species, characterized by the cylindrical or obovate-oblong, brown, fasciculate sporangia, fre- quently exhibiting metallic tints, and the dense orange-brown capillitium of spinulose threads. The spores vary in marking when magnified 1200 diameters from being almost perfectly smooth, to minutely but distinctly warted. On rotten wood, moss, &c. Britain (Apetliorpe, Weybridge, Twycross, Birmingham, Bulmer, N. Yorks ; Orton Wood, Leicester; Wothorpe, Scarboro', Carlisle); France; Germany; 160 A Monograph of the Mijxogastres. Switzerland; Italy; Belgium; Denmark; Hungary; Finland; Bohemia; Venezuela; United States; Cuba; Ceylon; Australia; New Zealand. (Rostafinski's Synonyms.) Clathroidcs 2-)yriformc, Hall, t. 1, f. 5 (1742). TricMa, Hall, t. 48, f. 5, No. 2167 (1798). Lycoperdon vesjparium, Batsch, t. 30, f. 172 (1786). Stemonitis ciimabarina, Roth. Fl. Germ., 347 (1788). Lycoxjcrdon favaceum, Schr. Fl. Bav., ii., 667 (1789). TricMa 2njriformis, Hoffm. V. Cr., t. 1, f. 1, (1790). Stemonitis fasciculata, Pers. in Gmel. Sys., 1468 (1791). Stemonitis vesparia, Gmel. Sys., 1070 (1791). Trichia rubiformis, Pers. Disp., t. 1, f. 3, t. 4, f. 3 (1797); Berk. Ann. N. H., No. 218; Cooke, Hdbk., No. 1177. Trichia rubiformis, /3 minor, Pers. Disp., 54 (1797). Lycoi^erdon ferrugincum, Hedw., t. 10, f. 1 — 4 (1802). Trichia chalybea, Chev. Fl. Par., t. 9, f. 24 (1827). Trichia Ncesiana, Corda Ic, i., f. 288 c (1837). Trichia Ayresii, B. and Br., Ann. N. H., No. 390 ; Cooke, Hdbk., No. 1179. Arcyria paradoxa, Mass. (fig. 272). Sporangia scattered or aggregated, sessile on a broad base, hemispherical or irregularly elongated, smooth, rather shining, dirty ochraceous, iomer surface of the wall with a layer of organic particles ; mass of capillitium and spores pale lemon-yellow ; capillitiura scanty, threads 4 — 5 /^ thick, much contorted and forming a very loose net with hut few ahrniot free tips, spirals very much crowded, not prominent ; spores globose, very minutely loarted, 8 — 10 /x diameter. Hemiarcyria paradoxa, Mass., Rev. Trich., p. 350, 357, fig. 35. On wood. Weybridge ; Surrey. Sporangia when hemispherical about '5 mm. diameter, 1 — 2 mm. when elongated. Distinguished amongst species with Arcj/ria. 161 wartcd spores by the densely crowded spirals of the elaters, and the presence of organic particles on the inner surface of the sporangial wall. A note by Currey, who collected the specimen, now in the Currey collection in the Kew herbarium, says, "The spores of this specimen sown in water produced de Bary's zoospores in 24 hours." Arcyria Bucknalli, Mass. (figs. 270, 271). Sporangia sessile on a broad or narrow base, seated on a very thin hypothallus, circular, reniform, or subangular from mutual pressure, wall very thin, gilvo-ochraccmis, soon disappearing; mass of spores dull orange ; capillitium well-developed, threads sparingly branched and combined to form an irregular open network with numerous, long, free tips, 4 — 5 n thick, walls with annular ridges mostly crowded, but here and there scattered, and sometimes passing into a spiral, the ridges with numerous thin, straight spines 3 — 4 jj. long the free tips irregularly swollen, ivithout spirals or rings, hut h^istling with s^yincs, as arc also certain interstitial sioollen portions ; spores globose, pale yellow, minutely wartcd, 7 — 9 ju diameter. Hemiarcyria Buchiallii, Mass., Grev,, v., 18, p. 27 (1889) ; Bucknall, Fungi of the Bristol district, in Bristol Nat. Soc. Proc, vol. vi., pt. II., 1889, 1890, p. 4, pi. 2, f. 5. On wood. Bristol. Generally crowded, about o mm. diameter, but extending to 1*5 mm. when isolated and elongated. Most closely allied to Hemiarcyria Wigandi, Rost., but at once distinguished by the larger sporangia, the markings on the elaters being mostly rings and not spirals, and in being furnished with numerous spines. The present species combines the characters of Rosta- finski's genera Hemiarcyria and Cormcvia, the spirals on the threads of the capillitium pointing to the former, and the numerous free arms to the latter. The spirals technically separate this species from 02)hiothcca as understood in the present work. 162 A Monogmpli of the Myxogastrea. Arcyria melanopeziza, Mass. Sporangia sessile, creeping, suhierctc, generally forming rings, 1 — 2 mm. long, very Hack, scarcely or not at all shining, very smooth; wall black, opaque, subcellular, rather coriaceous, splitting longitudinally and deliiscing in a valvate manner; capillitium yellow, protruded elastically, threads round, 4 — 5 ix thick, combined into a loose net, everywhere covered with erect spines, 5 — 6 x 1 /a, spirals almost obsolete ; spores elliptico-glohose, pap)illoso-scalrid, 10 — 12 x 10 /x, yellow. Hemitriehia melanopeziza, Spegazzini, Fung. Arg., Pug. IV., n. 268. Hemiarcyria melanopeziza, Berlese, Sacc, SylL, vii., 1, n. 1520 ; Mass., Rev. Trich., p. 356. On bark. Brazil. Looking exactly like some black Peziza. Arcyria calyculata, Mass. Sporangia simple, gregarious, stipitate, globose or elliptical, dirty foxy-brown, 1 — 2 mm. diameter; stem 3 — 5 mm. long, 200 — 250 /a thick, round, glabrous, rather tough, apex dilated into a little dimidiate cup equal to the pcridium, base dilated, fibrillose, colour of the sporangium ; spores and capillitium dingy yellowish -fulvous; elaters 7 — 8 }x thick, cylindrical, yellowish, sparsely branched, free tips acute ; spirals 3 — 5, flat, not very conspicuous, separated by interspaces their own width, spinulose ; spores discoideo-lenticular, concavo-convex, margin muriculate, 10 X 3 fA diameter. Hemiarcyria calyculata, Speg., Fung. Arg., Pug. III., n. 92 ; Sacc, Syll, n. 1518; Mass., Revis. Trich., p. 356. On rotten willow trunk. Argentine Republic. It is very doubtful as to whether the spores in the normal condition present the shape described by Spegazzini ; the spores of many species become concavo-convex when dry, but if soaked for a sufficient length of time, assume the usual spherical form. Arri/rlti. if]3 Arcyria stipitata, Mass. Sporangia pyriform, from 2—5 on a common stem, or solitary, pale kmon-yellou; opaque; stem elongated, equal, dark hrown hlack, longitudinally rugulose; mass of capillitium and spores dingy ochre ; capillitium dense, much hranched and forming a net without free tips, threads 4—5 // thick, s^nrals very open, rather distant, thin, not prominent; spores globose, minutely warted, 7 — 8 )a diameter. a. genuina. Sporangia single on an elongated stem. 13. fascieulata. Sporangia fasciculate on a common stem. Eemiarcyria stipitata, Mass., Rev. Trich., p. 354, fig. 32. On palm stems. Java. (Type in Herb. Berk., Kew.) Scattered or aggregated, 3—4 mm. high, stem about 2 mm. long, thin, hollow. Capillitium elastic, protruding after dehis- cence. Allied to Arcyria clavata, but distinct in the dense capillitium without free tips, the loose spirals, and the long, thin, black stem. Arcyria Wigandii, Mass. Sporangia clavate, discoid, or irregularly subrotund, very small, almost sessile, mass of spores and capillitium hay or flcsh-eolcur, verging on yellow ; elaters rarely branching, spirals one or two, flcrnous, either separated hy interspaces from three to four times their oivn width, or croivded and almost forming rings, tips scarcely narrowed, truncate and inflated ; spores minutely verruculosc, 10 — 11 ju, diameter. Eemiarcyria Wigandii, Rost., Mon., p. 2(i7, fio-. 232 ; Sacc, Syll., vii., 1, no. 1517; Cooke, Myx. Brit., fig. 232; Mass., Rev. Trich., p. 356. Germany. (Rostafinski's Synonym.) Trichia ahietina, Wigd., 1, c, p. 33, t. 2, f 11 (18(j3). 164 A Monograph of the Mi/ocoijaiftres, § § Simres with raised hamis combined to form a network. Arcjrria chrysospora, Mass. (figs. 268, 269), Sporangia sessile on a broad base, generally closely aggre- gated, bright ochraceous yellow ; mass of capillitiura and spores yellow ; threads 5 /x thick, forming a loose net with many free ends, which generally terminate in slightly expanded, smooth, bent or straight conical apices, i^/pircds four, rather close, not ^prominent, connected by less prominent ridges rimning paixdlel to the long axis of the thread ; spores globose, with raised fiat bands forming a polygonal network, 16 /x diameter. Hemiarcyria chrysospora, Lister, Grev., vol. v., p. 126; Mass., Rev. Trieh., p. 357, fig. 37- On larch twigs lying on the ground and on the surrounding herbage. Lyme Regis. A fine species with the sporangia reaching to 1 mm. diameter, approaching in habit and general structure the form sessilis of Arcyria scrpula, but quite distinct in the larger spores, the thinner capillitium threads without spines, and having the spirals connected by ridges parallel to the axis of the elaters. From three to five polygons on a hemisphere of the spore. Arcyria serpula, Mass. (figs. 273, 274a). Either a vein-like, creeping plasm bdiocarp, usually anasto- mosing to form a net, or subglobose, scattered, and sessile on a broad base, wall thin, fragile, yellow, sometimes tinged brown ; mass of spores and capillitium yellow or orange ; threads of the dense capillitium 5 — 6 jx thick, forming a net with numerous free ends which usually terminate in a smooth, thin, tapering spine 8 — 10 /x long, sjnrals thin, not prominent, rather distant, furnished loitli numerous long, slender spinules ; spores globose, with narroio, raised fiat bands forming an irregular network, 10 — 12 [X. diameter. a. plasmodiocarpa. Plasraodiocarp vein-like, usually forming a network. Arcpria. 105 /3. sessihs. Sijorangia sessile on a broad base, scattered or gregarious, capillitium orange, elastic. Ilemiarcyria serpula, Host,, Mon., p. 267, lifs. 200 227 228 • Cooke, Myx. Brit, p. 68, figs. 200, 227, 228 ; Schroeterp' 115 ;' Sacc, Syll., vii., pt. I., n. 1514; Raiink., Myx. Dan., p. 04, t. 8,' f. 16 ; Mass, Rev. TricL, p. 357, f. 34. Exsicc.—FMcke\, Fung. Rhen., 2692 (as HemitricMa contorta (Ditnj.) Rost.). On rotten wood, branches, leaves, &c. Britain (Carlisle; specimen in Herb. Berk., from Sowerby's herbarium, no locality' called To^ichia reticulata, undoubtedly British); Germany;' Sweden; Belgium; Italy; United States; Cuba; St. Vincent,' Bombay; N. W. Australia; New Zealand; Ceylon. Distinguished amongst the clear yellow species by the very spiny elaters and the reticulate spores. The subglobose sessile form has not, so far as I am aware, been met with in Britain, but, although differing in habit and in the bright orange elastic capillitium, agi-ees in the structure of the elaters and the spores. The type of the form sessilis is in the herbarium of the Paris Museum, and was collected in Cuba. (Rostafinski's Synonyms.) Mucoo^ serpula, Scop. Fl. Carn., t. 65 (1772). Lycoperdon lumhricak, Batsch., f. 174 (1786). Trichia spongioides, Vill. Fl. Dauph., 1061 (1789). Stemonitis hcmhricalis, Gmel. Sys., 1470 (1791). Trichia orticidata, Pers. Disp., 10 (1797) ; Ic. and Desc, 1. 12, f 1 Trichia serjmla, Pers. Disp, 10 (1797); Eng. Fl, v., 320; Cooke, Hdbk, 1189. Trichia serpula, /3 Sjwngioidcs, Pers. Syn., 181 (1801). Trichia vcnosa, Schum. Saell, 1456 (1803). Hyporhamma reticulatum, Corda Ic, v., 34 (1842). Trichia rctiformis, Payer. Crypt., I 574 (1850). Arcyria clavata, ^lass. (fig. 124). Sporangia varying from clavate to globose, ycUounsh, p)olishcd, 166 A Monograph of the Myxogastres. stem thin, length variable, yellow, often reddish at the base, filled with large, glolosc, or mlangular cells, ivliich jmss upwards into normcd spores; mass of spores clear yellow, ochraceous- orange, or tinged with olive ; threads of capillitium 4 — 5 fx thick, forked repeatedly and combined into an irregular open net, free ends not mcmeo'ous, obtuse, sometimes sligldly swollen, spirals thin, not prominent, rather distant ; spores globose, very mimdely and irregularly reticidated, 8 — 10 fx diameter. Hcmiarcyria clavata, Rost., Mon., p. 264, figs. 205, 207, 210, 238; Cke., Myx. Brit., p. 68, figs. 205, 207, 210, 238; Sacc, Syll, vii., 1, n. 1513; Mass., Mon. Trich., p. 31 ; Raunk., Myx. Dan., p. 64; Schroeter, p. 114. Exsiee. — Fckl., F. Rhen., 1434; Jack, Leiner u. Sitzenb, Krypt. Badens., 621 ; Ellis, N. Amer. Fung., 523. On decayed wood, &c. Britain (King's Cliffe, Apethorpe, Scarboro', Carlisle) ; France ; Germany ; Denmark ; United States ; Cuba ; Brazil ; Ceylon ; Benin Islands. Scattered or gregarious, 1'5 — 2 mm. high, stem slender, often attenuated downwards and longitudinally wrinkled, sometimes very short, especially when the sporangium is almost globose. The spores are usually covered with very thin, slightly raised ridges combined into a very fine, irregular network, but the ridires are sometimes short and distinct or even wart-like. (Rostafinski's Synonyms.) Clathrus 2Jedcdits, Schm. Ic, t. 33, f. 1, 17 (1776). Sphaerocarj)US iiyriformis, Bull., t. 417, f. 2 (1791). Stemonitis pyriformis, Gmel. Sys., 1469 (1791). TricJda pyri/ormis, Sibth. Fl. Ox., 406 (1794); Sow., 400, f. 6. Triehia clavata, Pers. Disp., p. 11 (1797) ; Eng. Fl., v., 320 ; Cooke, Hdbk., 1183. Ti'ichia citrina, Schum. Saell., 1460 (1803). Arcyria trichioidcs, Rudolph, Linnaea, p. 120 (1829). Triehia erythropiis, Borszczow (1856). Triehia oUusa, Wigand, p 30, t. 11, f. 4 (1863). Triehia Thwaitcsii, B. and Br., Ceylon Fung., No. 776 (1873). Arcyria. 1G7 § § § ^^l^ores smooth. Arcyrio leiocarpa, Mass. Sporangia scattered or aggregated, obovate or pyriform, rarely almost globose, pallid, stem same colour, as long as diameter of sporangium; mass of capillitium and spores concolorous, or with a slight ochraceous tint; capillitium sparse, forming a loose net, threads 5 /x thick, spirals tliin, rather close, slightly pro- minent on the convex side of the bent threads, usually furnished with scattered, rudimentary spinules, free tips very rare or absent ; spores globose, smooth, 12 — 14 ju diameter. Hemiarcyria leiocarpa, Cooke, Myx., U. States, in Ann. Syc. Nat. Hist., New York, xi., n. 12, p. 405 ; Cooke, Myx. Brit., figs. 252, 255; Sacc, Syll., vii., 1, n. 1519; Mass., Rev. Trich., p. 353, fig. 33. In Saccardo's Sylloge VIT., pt. I., p. 440, n. 1519, Rostafinski is given as the author of the present species, and the reader is referred to "Rost., Mon., p. 267" for the description, but in reality the jDresent species is not included in Rostafinski's work. Arcyria leocarpoides, Mass. Sporangia globose or pyriform, 0'6 — 0'8 mm. diameter, flesh- red or fulvous, often depressed and deeper coloured at the apex, not at all or only a little shining, very smooth, cortex rather thick, subcartilaginous, soon breaking away in pieces above, usually persistent, cup-like at the base ; stem rather rigid, erect, brownish or blackish, slender, even or subrugulose, about as long as diameter of sporangium ; capiUitimii after dehiscence protruding elastically, attached at the base, very persistent, brownish-fulvous, or olive-fulvous, threads slender, cylindrical, 5 — 6 ii thick, foi-ming a dense net with numerous terete, aWuptly rou7ided free arms, threads with 3 — 4 spirals, very smooth, not papillate; spores globose, 8 — 10 )a diameter, very smooth, proto- plasm granular, fulvous-olive. Cornuvia leocarpoides, Speg., Fung. Arg. Pug. I., n. 264 ; Sacc, Syll., vii., 1, n. 1546. On rotten wood. Brazil. 168 A Monograph of the Mifjcogastres. Tbe stipitate sporangia and elastically protruded capillitium attached at the base, as also the spiral markings on the threads prove the present species to be an Arcyria, in which genus free arms to the capillitium are frequent in several species. Arcyria Karsteni, Rost. Sporangia effused, vermiform, sinuous, sometimes forming irregular reticulations, or hemispherical, scattered, and sessile on a broad base, varying from dirty ochraceous brown to dark chestnut ; mass of capillitium and spores dingy ochre ; threads often irregularly branched, and forming a very loose net, 3 — 4 \x thick, witli scattered, inflated portions 12 — 15 ix tldch, and 30 — 50 /x long, spirals very indistinct, a few scattered rudimentary spines now and then present, free tips not distinctly attenuated, usually abrupt or clavate; spores globose, smooth, 12 — 12 jot diameter. \ Hcmiarcyria Karsteni, Rost., Mon. SuppL, p. 41 ; Karst., Myx. Fenn., iv., p. 14^; Schroeter, p. 115; Sacc, Syll., vii., 1, n. 151G ; Mass., Rev. Trich., p. 342, fig. 8G. (Specimen from Ceylon in Herb. Berk., named by Rostafinski.) On wood. Finland ; Silesia ; Ceylon. Characterized by the threads of the scanty capillitium being considerably swollen at intervals, and by the indistinct spirals. Arcyria pusilla, Mass. Sporangia rather closely gregarious, subcylindrico-elliptical, 0'4 — 0"5 mm. high, 0*15 — 025 mm. diameter, rather obtuse above, abruptly subtruncate below, with only a trace of a stem or altogether without; at first hlood-red with an amber tinge, afterwards rose-colour ; capillitium rather dense, forming a rose- coloured network, threads round, 3 — 4 ix thick, spirals three or four, furnished with minute spinules ; spores rose or flesh-colour, globose, smooth, 7 — 9 ju diameter. Hemitriehia p)usiUa, Speg., Fung. Arg. Pug., IV., n. 269. Hcmiarcyria j^ws27/«, Berlese, Sacc, Syll., IV., 1, n. 1512; Mass., Rev. Trich., p. 352. On bark. Argentine Republic. Ai'ci/ria. ig9 Arcyria fuliginea, Cke. and Mass. (fig. 113). Sporangia (?), mass of capillitiuni and spores smolcy-olive, threads round, 6 — 7 /^ diameter, thivldy and equally covered ivith slender s^nnes 3 — 4 /x long, combined to form a dense, irreo"ular network ; spores globose, smooth, 8 /a diameter. Hemiarcijria fuliginea, Cke. and Mass., Grev., vol. xvi., p. 74. On living leaves. Mount Wilson, N. S. Wales. Nothing is known of the sporangium, which had been com- pletely crushed and lost; capillitiura forming a large mass spreading over the leaves. May possibly prove to be a broadly effused, plasmodiocarp form of an 0])hiotheca. Sub-sect. Tricheae. Oligonema, Host. Wall of sporangium single, dehiscing irregularly; capillitium scanty, composed of free, simple or branched elaters, furnished with ring-like thickenings, or a single very diffuse, rudimentary spiral, tips obtuse. Oligonema, Rost., Mon., p. 2.91; Cooke, Myx. Brit., p. 77; Sacc, Syll., p. 436 ; Mass., Rev. Trich., p. 21. Agreeing with the genus Trichia in having the elaters per- fectly free, but readily distinguished by the rudimentary mark- ings on their walls, never having more than a single, indistinct, very open spiral, which may be present on one portion of an elater and absent on another part, or not unfrequently altogether absent from all the elaters of one sporangium, and present on some of those from another sporangium taken from the same group. A second type of ornamentation on the walls of the elaters iii the present genus consists of annular or ring-like thickenings, which present the appearance of thin, flat discs, rather larger than the diameter of the elater, and placed at right angles to its long axis. These ring-like thicken- ings are very unequally distributed, five or six being sometimes met with at unequal distances on an elater, whereas tliev mav 170 A Monograph of the Myxogastres'. be entirely absent from all the elaters next examined. These annular thickenings are also met with on the threads of the capillitium of some species of Cornuvia, Didymium, Tilmadoche, &c. In rare instances the two ends of an elater coalesce and form a closed ring, as figured by Rostafinski in Oligonema nitens, Rost., Mon., fig. 198. Distrih. Europe ; North Africa ; North America. Species 7. * Epispore reticulated. Oligonema nitens, Rost. (figs. 193 — 195). Sporangia densely crowded, in one or several superposed layers, when scattered, sessile on a broad or slightly contracted base, clear jprimrosc-ycllow, very smooth and shining ; mass of capillitium and spores yellow; elaters scanty, 4 — 5 /x thick, simple or sparingly branched, very variable in marking, with scattered, thick, projecting rings, or a very indistinct, open spiral on the whole or part of an elater, or a combination of both, not unfrequently perfectly smooth, tips obtuse or more or less pointed ; spores globose, with narroio raised ridges of varying thickness, forming an irregular network, 11 — 13 fi diameter. Oligonema nitens, Rost., Mon., p. 291, f. 198 ; Cke., Myx. Brit., f. 198 ; Schroeter, p. 108 ; Sacc, Syll., n. 1488 (the sporangium described as of a gilvous colour by mistake) ; Mass., Rev. Trich., p. 347, f. 29. Oligonema havarica, Balf. and Berl., Sacc, Syll, n. 1490. Oligonema nitens, a havarica, Mass., Rev. Trich., p. 347, f. 25, Trichia nitens, Libert, PL Cr. Ard., fasc. iii., n. 227. Exsicc. — Lib., PI. Crypt. Ard., fasc. iii., n. 227 (as Trichia nitens) ; Klotzsch, Herb. Myc, (Rabeuh.) 137 (as Trichia circum- scissa^ ; Fuckel, F. Rhen., 2198 (as Trichia nitens) ; de Thum., Myc. Univ., nos. 399 and 1497 (as Trichia Bavarica). On wood, bark, &c. Britain (Carlisle, Bristol, Birmingham) ; France ; Bavaria ; Germany. Sporangia "5 — 1 mm. diameter, clear yellow, polished ; the elaters are extremely variable in marking, as is also the Oligonema. 171 epispore, but the latter always in the markings forming a reticulation. Oligonema flavidum, Mass, Sporangia crowded, sessile, Iriglit yclloio, shining, mass of capillitium and spores bright yellow; elaters scanty, short, 4 — 5 1^ thick, sometimes with an elliptical interstitial swelling, tips variable, subacute or inflated, densely covered with minute raised jjoints ; spores globose, with thin raised hands combined to form a very irregidar nctioorh, bands not punctate, 7 — 9 fx diameter. Ferichaena fiavida, Peck, 26th Annual Report of New York State Mus., p. 76 (1874). On moss, wood, &c. United States. The sporangia are irregular in form, circular, angular, or elongated. Forming patches of considerable extent. Superficially resembling 0. nitens, but distinct in the elaters and spores. The elaters agree with those of 0. minntula in being rugulose or covered with small points or band-like wrinkles, but in the present species the spores are smaller and the reticulations more irregular in form and size. Oligonema minutulum, Mass. Sporangia scattered, rarely aggregated, sessile on a narrow base, lemon-yelloiv, dull, capillitium very scanty, elaters simple, short, cylindrical, 5 — C ix thick, ontgulose, and tvith a very open, indistinct spiral, tips obtuse, rounded ; spores globose, and with slightly raised, flattened bands forming a netivork of mcmerous, almost regular and eqital-sizcd polygons, 12 — 14 \x diameter. Oligonema minntula, Mass., Rev. Trich., p. 848, fig. 20. Type in Herb. Berk., Kew, no. 10902, marked " Trichia minutula, D. R., et Montag., Algiers," Algeria. So far as I am aware, the present species has not been described by Montague. Related to 0. nitens, but known by the scattered, dull sporangia, and the very few, short elaters having thick, rugulose walls, with an indistinct open spiral ; 172 A Monograj)h of the Mi/.rogastres. in rare instances a swollen portion 15 — 20 )u long, and 8 — 12 \i thick is present near the middle of an elater, but there is no indication of the narrow, ring-like thickenings as in 0. nitcns. From 7 — 9 complete polygons present on a hemisjDhere of a spore. * * Episporc warted. Oligonema Broomei, Mass. (figs. 185, 186). Sporangia scattered or in small grotq^s, adnate by a broad base, hemispherico-depressed, circular or irregularly elongated, wall smooth, dark hrown, dull ; mass of capillitium and spores reddish-ochre ; elaters cylindrical, 3 — 4 /x thick, irreff^ilarly hranched, dull-orange, furnished with narrow, ring-like thicken- ings, at regular distances, tips obtuse ; spores rather coarsely warted, dull orange or yellowish, globose, 13 — 14 ju diameter. Oligonema Broomei, Mass., Rev. Trich., Journ, Roy. Micr. Soc, 1889, p. 345, pi. vii., f. 22. (Type in Herb., Broome in Brit. Mus., marked " Trichia scrpula" (?).) On bark. Britain (Warleigh, Somerset). The present species is most neai'ly allied to 0. aeneum, Karst., which has up to the present been only met with in Finland, and is distinguished by the crowded habit, polished sporangia with metallic tints, and smaller spores. When circular the sporangia vary from "5 — 1 mm., and when elongated 2 — 3 mm. long. Oligonema aeneum, Karst. Sporangia densely crowded, often confluent and venulose, rarely scattered, orbicular or angular from mutual pressure, depressed, sinning, with copper, green, or olive metallic tints; threads of the capillitium free, 2 — 3 /a thick, with scattered thickenings in the form of circles; spores globose, wao'ted, rather ferruginous or pale reddish-ochre, 12 /a diameter. Oligonema aeneum, Karsten, Myc. Fenn., iv., p. 125; Sacc, Syll, n. 1487 ; Mass., Rev. Trich., p. 346. Finland. Olhjoneind. 173 Oligonema furcatum, Buckiuill. Sporangia scattered, globose, shining, lirujlit cJiromc-ycUoir, as well as the capillitium and spores; elatcrs cylindrical, simple or branched, slightly thickened at the obtuse ends, with a faint open spiral, 3 — 4 /x diameter ; spores globose, minatdy luartcd, 11 — 12 jM diameter. Oligoneii\a furcatum, Bucknall in litt. On a rotting trunk. Britain (Abbotts Leigh). Superficially closely resembling Oligonema nitens, from wliich however it is quite distinct in the minutely warted spores, and the absence of thickened rings on the usually furcate elaters. Oligonema brevifilum, Peck. Sporangia crowded in effused heaps, bright ochraceous yellow ; elaters few, short, cylindrical or suhfusiforiii ; spores globose, rugose, 11 ix diameter. Oligonema nitens, Peck, in 81st Report of State Agric. Mus., p. 42; Sacc, Syll., n. 1489; Mass., Rev. Trich., p. 346. On mosses. United States. Mr. Wingate of Philadelphia thinks that the present species will prove to be an abnormal condition of 0. fiavida. Trichia, Haller (emended). Wall of sporangium single, dehiscing irregularly ; capillitium consisting of free, simple or branched threads, having the wall furnished with raised bands arranged in a spiral manner ; spores globose, epispore smooth, or variously ornamented. Trichia, Haller, Helv., iii., p. 114 ; Rost., Mon., p. 243 ; Cooke, Myx. Brit., p. 61 (in part) ; Sacc, Syll., vol. vii., pt. I., p. 438 (in part) ; Mass., Mon. Trich., p. 328. The genus is readily distinguished by the presence of well- developed external ridges arranged in a spiral manner on the perfectly free elaters or threads of the caiiillitium. The elaters are in most species unbranched, cylindrical or fusiform, and more or less attenuated at the tii)s. In a few species the 174 A Monotjr(n>k of the MiixogaMres. elaters are branched, the free ends varying from three to ten in number. Tlie only other genus with free elaters is Oligonema, but here the spirals are at most rudimentary and the tips obtuse. Distrih. All temperate and tropical countries. Species 30. A. SiJorcs smooth. Trichia Carlyleana, Mass. (173—175). • Sporangia clavate or cylindric-oblong, stipitate, dark brown, often with a purple tinge, smooth ; stem usually about half as long as sporangium, equal, or slightly thickened downwards and expanded at the base into a small circular hypotiiallus ; inner surface of sporangial wall and holloiv of stem with numerous rather large organic lumjjs, of a bright reddish-purple colour; mass of spores bright yellow ; elaters fusiform, 5 — 6 \x at thickest part, simple or frequently branched, tips running into a long, smooth, very thin, straight or Hexuous spine, spirals crowded, thin, not prominent; spores globose, smooth, 10 — 12 /x diameter. Trichia Carlyleana, Mass., Mon. Trich., p. 5, f. 15. On wood. Britain (Carlisle). Sporangia in fascicles of 3 — 5 ; 2^3 mm. high. Superficially resembling some forms of Trichia fragilis, but quite distinct in the smooth spores, and the narrow, crowded, and not at all prominent spirals of the elaters. The colouring matter in the organic lumps on the wall of the sporangium and in the stem is soluble in dilute potassic or ammonia hydrate. Trichia heterotrichia, Balf. Sporangia sessile in clusters, dark yellow, wall thick, tough and leathery, inner layer areolate ; elaters few, cylindrical, 7 /x diameter (thickenings excluded), with walls of medium thick- ness, irregularly and variously thickened, either with spines often twice diameter of elaters, or with short prickles or warts, or with complete or half-rings, or sometimes with interrupted and irregular spirals leaving large intervening unthickened portions, swollen towards the extremities, and ending in a I'rlcliia. 175 tapered, rarely smooth, arcuate or twisted point, in lengtli twice the diameter of elater; spores globose, pale yellow, smooth, 16 — 17 [x diameter. Trichia heterotricJda, I. B. Balfour, Grev., vol. x,, p. 117; Sacc, SylL, n. 1505; Mass., Mon. Trich., p. 5, f. 16. On bark. Britain (in Herb. Currey, Kew ; no locality). A species resembling most nearly forms of Trichia varia, Pers., but the few elaters with the very varying sculpturing and the larger smooth spores sufficiently separate tliem. (I. B. B.) B. S])orcs luarted. § Elaters fusiform. Trichia frag^ilis, E,ost. (figs. 7—10). Sporangia var3;ing from globose to pyriform, stipitate, solitary, or fasciculate on a common stem, wall smooth, blackish or purple-brown, pale brown or yellowish ; stem dark, longitudinally wrinkled, equal or tapering upwards, erect or curved, often twisted in the compound forms, inside of the wall often with coloured organic lumps ; mass of capillitium and spores varying from dull orange-brown to primrose-yellow, separated from the hollow of the stem by a membrane ; elaters fusiform, 4 — 5 y. at thickest part, sometimes branched, sinrals flat, rather broad, not very prominent, tips smooth, tapering to a thin point, spores globose, minutchj warted, 11 — 14 ju, diameter. a. genuinct. Sporangia pyriform, solitary or fasciculate, clear or black-brown, opaque ; mass of capillitium and spores varying from reddish-brown to dirty ochre ; stem erect. /3. Lorinscriana. Sporangia pyriform, solitary or fasciculate, reddish- brown, polished; mass of capillitium and spores dirty ochraceous ; stem generally curved. y. serotina. Sporangia clavate or pyriform, solitary or fascicu- late ; mass of spores and capillitium clear yellow to ochraceous ; stem erect or curved. 8. lateritia. Sporangia subglobose, solitary or fasciculate, almost black ; mass of capillitium and spores dark orange-brown ; stem attenuated upwards. 176 A Mono(jraph of the MiiJro(jastres. Trichia fra;/ilis, Rost., Mon., p. 246, figs. 203, 204, 225, 226 (in part); Cooke, Myx. Brit., p. 63, figs. 203, 204, 205, 226 (iu part) ; Sacc, Syll., vii., 1, n. 1494 (in part) ; Balf., Grev. v., 10, p. 116 (in part); Mass., Mon. Tricli., p. 6, figs. 14 and 28. Trichia latcritia, Lev., Ann. Sci. Nat., Ser. III., vol. v., p. 167; Sacc, Syll., vii., n. 1494 ; Balf., Grev., vol. x., p. 116. Trichia hotrytis, Schroeter, p. 112; Raunk., Myx. Dan., p. 67. Sphaerocarpus fragilis, Sow., t. 279. ^a-szcc— Cke., Fung. Brit, 612; Rab., Fung. Eur., 244; Jack, Leiner. u. Sitzenb. Krypt. Badens., 329; Erbar. Critt. Ital., 640; Ellis and Everh., N. Amer. Fung., Ser, II., 2097 and 2098 ; Fuckel, Fung. Rhen., 1437. On wood, twigs, &c. Britain (Brighton, Kew, Gloucester, Castle Howard, Yorks ; Carlisle, Appin, N. B.) ; Europe ; United States ; Canada ; Chili ; S. Africa ; Ceylon ; S. W. Australia ; Tasmania ; N. Zealand. A variable species in colour of sporangium and mass of spores and capillitium. The distinguishing characters are the fusiform elaters with long, smooth, tapering tips and flat spirals, and the warted spores. Varying from 2 — 4 mm. high ; solitary, or in more or less confluent clusters of from 2 — 7 on a common stem, which is obviously composed of several confluent stems, and hence forming a stipitate aethalium. The elaters are sometimes branched near the tips. I have examined the type specimen of Trichia latcritia in the Paris Museum, and find that it agrees exactly with Trichia fragUis in the structure of the spores and capillitium. (Rostafinski's Synonyms.) Lycoperdon hombaciwwni, Batsch, El., p. 153 (1783). Stemonitis hotrytis, Pers. in Gmel. Syst., 1468 (1791). Trichia hotrytis, Pers. Disp., p. 9 (1797); Ic. Pict., t. 12, f. 1—2. Trichia hotrytis, 13. minor, Pers. Disp., 54 (1797). Trichia serotina, Schrad., Journ., t. 3, f. 1 (1799) ; Engl. FL, v., p. 310; Cooke, Hdbk., No. 1181. Trie hid. 177 SphaerocarjMS fragilis, Sow., t. 279 (1803). TricMa notata, Fl. Dan., 1680 (1823). Trichia hadia, Fr. Stirp. Femsj., 83 (1825). TricJiia pyriformis, Fr., S. M., iii., 184 (1829) ; Curr., Micr. Journ., v., p. 129; Cke., Hdbk., n. 1178. Tridda Zorinseriana, Corda, Ic, f. 228 D (1837) ; Curr., Micr. Journ., v., p. 129 ; Cke., Hdbk., n. 1180. Trichia pyo'iformis, (3. scrotina, Rost., in Fckl. Synib., 2 N., 75 (1873). Craterium floriformc, Schu., Ann., No. 2307. Al'wisia homharda, B. and Br., Ceylon Fung., No. 784, t. 2, f. 6 (1873). Trichia purpurascens, Nyl. (figs. 183, 184). Sporangia stipitate, ovate or spherieo-ovate, solitary or gre- garious, jmrjjlish-rcd, opaque; stem longitudinally wrinkled, erect or cernuous, rather firm and thickish, expanding at the base into a small hypothallus, coloured like the sporangium, which it equals or exceeds in length ; mass of elaters and spores bright ochraceous ; elaters rather short, fusiform, attamatcd into long, tapering, smooth, straight or fiexuoiis tips, simple or branched, about 5 fl thick at the centre, spirals sharp-edged, rather ^jro- minent and distant ; spores globose, verruculose, yellow, 9 — 11 ft diameter. Trichia purpurascens, Nyl., in Siillsk. pro Faun, et Flor. Fenn. notis. Ny., Ser. 11., 2, p. 12G ; Mass., Rev. Trich., p. 332; Sacc, Syll., no. 1508; Grev., v., 18, p. 27 (1889); Karst., Myx. Fenn., iv., p. 137. On fir-wood, moss, &c. Britain (Carlisle) ; Finland. Requires to be carefully distinguished from some forms of Trichia fragilis, to which species the present is closely allied. In T. purpurascens the spirals are sharp-edged, in T. fragilis flattened. The inner surface of the sporangial wall is studded with amorphous, organic purple particles ; the colouring matter is soluble in potassic or ammonic hydrate. 178 A Monograj^h of the Myxogastres. § § Maters cylindrical. * Sjnrals not spinulose. Trichia varia, Rost. (figs. 169—172). Sporangia scattered or aggregated, stipitate, or sessile on a narrow or broad base, turbinate or subglobose, smooth, yellow, dirty ochraceous or tinged with olive; stem when present blackish ; mass of capillitium and spores yellow ; elatcrs cylin- drical, 4 — 5 /x thick, spirals distant, prominent, more especially on the convex side when curved, rarely branched, tips smooth, tapering, straight or bent, 8 — 10 ju. long, sometimes shorter; the elaters are sometimes swollen at the commencement of the taper- ing tips; spores globose, minutely loarted, 10 — 14 /x diameter. Triehia varia, Host., Mon., p. 251, figs. 191, 202, 208, 212, 218, 237; Cooke, Myx. Brit., p. 63, figs. 191, 202, 208, 212, 218, 237; Schroeter, p. 112; Sacc, Syll., vii., 1, no, 1497; Raunk., Myx. Dan., p. 65, t. 3, f. 14, t. 4, f. 3; Mass., Mon. Trich., p. 9, f. 17. Exsicc. — Jack, Leiner u. Sitz., 419 ; Karst., Fung. Fenn., 288 and 690; Fuckel, Fung. Ehen., 1431 and 1433; Roum., Fung. Gal]., 1101 and cent. xiv. 1315; Rab., Fung. Eur., 799, 2137 and 2138 ; Thum., Myc. Univ., 1999 ; Sacc, Myc. Van., 794. On bark, wood, moss, &c. Britain (Kew, Bishop's Wood, Highgate ; Weybridge, Scarboro', Carlisle, Abergavenny, Appin, N. B.) ; France ; Denmark ; Germany ; Finland ; Italy ; Bo- hemia ; United States ; Tasmania ; New Zealand ; Australia. A variable species, sometimes with a distinct, dark-coloured stem, at others quite sessile on a narrow or broad base. Characterized by the cylindrical elaters with distant, prominent spirals, and the minutely warted spores. In the specimen in Rabenh., Fung. Eur., n. 2137, the warts show a slight tendency to become elongated and flattened. (Rostafinski's Synonyms.) a. Trichia varia, v. nigripes. Mucilago minima, Mich., t. 96, f. 4 (1729). Tric/n 179 3mholus albissimus, Hall, Herb., p. 8 (1742). Emholus, Hall, No. 2138 (1768). Mucor pyriformis, Scop. Fl. Carn., 402 (1772). Mucor lyomiformis, Leers Fl. Herb., IISO (1775). Mticor ladeus, Leers Fl. Herb., 1132 (1775). Stemonitis pyriformis, Willd. Fl. Ber., 409 (1787). Emholus ladeus, HofF. Veg. Cr., t. 2, f. 3 (1790). ^phaerocarpus chrysospermus, Bull., t. 417, f. 4 (?). Trichia olivacea, Pers. Obs., i., 62 (1796). Arcyria olivacea, Ransch (1797). Trichia cylindrica, Pers. Obs., ii,, 33 (1799). Trichia cm-data, Pers. Obs., ii.,- 33 (1799). Trichia nigripes, Pers. Syn., 178 (1801). a pyriformis, /3 cordata,y cylindrical, h vulgaris; Fl. Dan., t. 1313, f. 2; Curr., Micr. Journ., v., p. 128; Cooke, Hdbk No. 1185. Trichia craterioides, Corda, Ic, ii., f. 85 (1838). y Trichia varia, v. genuina. Lycogala luteum, Mich., t. 95, f. 4 (1729). Mucor quintus, Schff., 296 (1770). Mucor granulatus, Schff., No. 286 (1770). Lycoperdon vcsiculosum, Batsch., 283 (1786). Sphaerocarpus chrysospermus, Bull., t. 417, f. 4 (?). Stemonitis varia, Pers. in Gmei. Sys., 1470 (1791). Stemonitis vesiculosa, Gmel. Sys., 1470 (1791). Trichia olivacea, Pers. Obs., 1., 62 (1796). Trichia varia, Pers. Disp., p. 10 (1797) ; Eng. FL, v., 320 • Cooke, Hdbk., No. 1188. > > - , Lycojicrdoii luridum, Hedw. Obs., t. xi. a (1802). Trichia favoginca, Schum. Saell., 1455 (1803). Trichia applanata, Hedw., in DC. Organ, t. 60, f. 1 (1827). Trichia nitens, Fries. Sporangia sessile on a broad base, crowded, hemispherical, convex above, circular or angular from mutual pressure, jpnm- 180 A MonoffJ'aph of the Myocogastres. rosc-yeUmo, j^oli^hcd, shining; mass of spores and elaters dull- orange; clatcrs cylindrical, 14 — 16 /x thick, rather short, ending in a very short, abrupt, smooth apiculus, spirals rather pro- minent, distant, without spines; spores globose, with rather large, rounded warts, 14 — 16 ix diameter. Trichia nitens, Fries, MS.; Mass., Mon. Trich., p. 833, fig. 11. (Specimen from Fries in Herb. Berk., Kew, and named by Fries " Trichia nitens, Fr.") On wood. Upsala. A very fine large species, forming dense clusters 1 in. and more across. Externally resembling Oligoncma nitens, Rost,, distinct from T. varia in the polished, shining sporangia, and the thicker elaters, with very short, abruptly apiculate tips. Trichia inconsplcua, Rost. Sporangia very minute, Irovm, shining, subspherical, collected in clusters or scattered ; hypothallus absent ; elaters cylindrical, 3 '3 \x thick, tips pointed, 6 — 7 i^- long, curved, sometimes with elongated swellings near the ends, spircds 3 — 4, hut slightly 2')rominent, rather close; spores delicately vcro^uculose, 10 — 12 /x diameter. Trichia inconspicua, Rost., Mon., p. 259; Sacc, SylL, n. 1502 ; Mass., Mon. Trich., p. 335. Germany ; France. Trichia proximella, Karst. Sporangia substipitate or sessile, circular or irregularly angular from pressure, yellow or dingy ochraceous, polished ; mass of elaters and spores ochraceous; elaters cylindrical, sometimes branched, ending abruptly in a short, smooth, straight or oblique apiculus, 4 — 5 fx thick, sjjirals prominent, distant ; spores globose, rather coarsely wartecl, 12 — 14 /a diameter. Trichia proximella, Karst., Myc. Fenn., iv., p. 139; Sacc, Syll., vii., 1, no. 1507; Mass., Mon. Trich., p. 335. On wood. Finland. TricJiia. 181 Allied to Trichia nitens, but known by tlui much thinner elaters. Forming crowded j)atches of considerable extent. Trichia advenula, Mass. (figs. 142 — 144). Sessile on a broad base, densely crowded or with a few out- side scattered sporangia, circular, or subangular from mutual pressure, rather depressed, ^jWrnrose-^/cZ/o^y, shining; mass of capillitium and spores orange ; elaters cylindrical, 4 — 5 ju, thick, ustcally inflated at one or both ends, and also with' 1 — 3 interstitial swollen portions, the swollen ends terminating in a thin, straight, or usually Jiexuous slender spine, 15 — 20 /x long, spirals very close, but little prominent, almost obsolete on the swollen portions ; spores globose, minutely verruciUose, 12 — 14 fj, diameter. Trichia advenula, Mass., Rev. Trich., p. 336, fig. 38. On rotten wood. Glamis, N. B. Sporangia *5 mm. diameter, forming densely crowded patches 1 — 2 inches across. Most nearly related to Trichia minima, but distinguished by the long, slender tips to the elaters, and the interstitial swollen parts ; in T. minima the capillitium and spores are pale primrose in the mass, and not orange as in the present species. Trichia nana, Mass. Sporangia scattered or aggregated in small clusters, rarely exceeding "o mm. in diameter, sessile on a broad base, smooth, pale bright ochre, opaque, wall very thin ; mass of spores and elaters pale primrose yellow ; elaters cylindrical, short, 3 — 4 /x thick, spirals irregular, very distant, pirominent, tips abrupt, not apiculate, the spirals usually running quite to the end ; spores globose, minutely verruculosc, 6 — 8 [x diameter. Trichia nana, Mass., Mon Trich., p. 336. (Type in Herb., Kew.) On wood. Westbrook, Maine, U. S. Sporangia hemispherical or sausage-shaped ; by far the smallest of known species, resembling superficially T. minima, from which it differs in the distant and prominent spirals of 182 A Monograph of the Myxogastres. the elatcrs ; in the latter character it agrees with T. varia, but differs in the spirals not being markedly more prominent on the convex side of the bent elaters, the abrupt tips, and smaller size of every part. The elaters are rarely more than 200 ^ long. Trichia minima, Mass. Sporangia crowded, sessile on a broad base, circular, elliptical, or irregular from mutual pressure, _2-^a/c ^^'rimrosc-ydlow ; mass of elaters and spores same colour; elaters cylindrical, 5 — 7 \j. thick, ending in smooth, tapering points about 8 — 10 ^i long, spirals thin, rather distant, not prominent, without spines; spores globose, very minutely warted, 10 ft diameter. Trichia minima, Mass., Rev. Trich., p. 836, fig. 18. On wood. Britain (Oldham). Sporangia o mm. diameter, forming dense clusters. Allied to T. scahra, but distinct in the smaller size of every part, and in the absence of spines on the spirals of the elaters. Trichia contorta, Rost. Plasmodiocarp creeping, fiexuous, suhronq^orsscd, umler or hay- hrown ; mass of elaters and spores yellow ; elaters 2'5 — 3"5 /x, cylindrical, tips usually stvollcn and terminated by a long, slender spine; there is sometimes an interstitial swelling ; spirals indistinct ; spores globose, minutely warted, 12 — 15 ju diameter. Trichia contorta, Rost., Mon., p. 259, f. 229; Cooke, Myx. Brit., f. 229; Schroeter, p. 113; Sacc, SylL, vol. vii., pt. I., no. 1503; Raunk., Myx. Dan., p. 68, t. 3, £ 13; Mass., Rev. Trich., p. 337. On rotten wood. Britain ; Germany ; France ; Denmark ; Sweden; Australia, I have seen no specimen of the present species, which is said to have occurred in Britam. The cylindrico-compressed flexuous plasmodiocarp of a dark brown colour, and the elaters with long, spine-like tips and indistinct spirals mark the species. The elaters appear to agree in structure with those of Trichia Trichia. 183 advcnula, but the latter differs in every other character. The figure of the elater is copied from Rostafinski. (Rostafinski's Synonyms.) Lycogala contortum, Dit., in Sturm. Dent. Cr. Fl, t. 5 (1817). Trichia retimlata, b., Grev. Sc. Cr. Fl., t. 266 (1827). Ferichaena contorta, Fr., S. M., iii., 192 (1829). Licea contorta, Wallr., Fl. Cr. Germ., n. 2110 (1833). Hcmitricliia contorta, Rost., ap Fckl. Syn., 2, Nach., p. 75 (1873). Trichia subfusca, Rex. Sporangia stipitate, simple, very rarely double or triple, generally exactly spherical, exceptionally globose-turbinate, about ^ — I of a mm. in diameter. Colour of sporangia, a dull tawny brown above, shading to a dark brown at the base. Stipes uniform in diameter, and equal in height to the diameter of the sporangium, brown or brownish-black in colour, longi- tudinally rugose and separated from the cavity of the sporangia by the internal layer of the sporangium wall. Spores and capillitium concolourous, of a bright yellow colour in mass. Elaters cyhndrical 3'5 — 4 /u, in diameter, terminating generally in a smooth elongated end 10 — 12 /x long, which is either sharp or blunt, straight or curved to one side. Spirals four in number, non-spinulose, winding more or less unevenly, with interspaces as wide or wider than the thickness of the spirals. Spores 11"5 — 12*5 /ia in diameter, delicately but distinctly warted. Adirondack Mts., New York. This Trichia is more nearly allied to some of the forms of Tr. fragilis than to any other species. There seem, however, to be sufficiently well-marked specific differences. In addition to the different external characters, the elaters are undoubtedly cylindrical in a majority of the specimens. Occasionally individual sporangia are found in which the elaters are a little thicker in the centre, narrowing slightly toward the ends, but even these exceptions terminate abruptly in the same form of ends as the true cylindrical elaters. Occasionally also, as with 184 A Monogmi^ of the Myxogastres. all Trichias, sporangia will be found in which the elaters are branched or distorted, or have a tendency to bulbous expansions near the ends or in their course, but these are abnormal and exceptional. The specimens distributed under this number (2495) were collected in August 1889, with a few exceptions. The unusually wet season had the effect of rendering many of them much darker in colour than is indicated in the foregoing specific diagnosis, which was drawn from types developed and collected under the most favourable conditions. (Rex.) Trichia mhfusca, Rex, Proc. Acad. Sci. Phil., 1890, p. 192. Uxsicc. — Ellis and Everh., N. Amer. Fung., n. 2495. Trichia reniformis, Peck. Sporangia gregarious or clustered, sessile, subgiobose or reni- form, small, brown ; flocci few, short, sparingly branched ; spores globose, minutely echinulate, yellow-ochre, sometimes tinged with green, -0005 in. in diameter ( = 12 — 13 //). Peck, 26th Report of the State Museum, New York, p. 76 ; Sacc, SylL, no. 1510; Mass., Rev. Trich., p. 337. United States. I am not acquainted with the present species, and the inform- ation given by Peck is too scanty to insure future identification. § § Sjnrals sjmmlose. Trichia erecta, Rex. Sporangia stipitate, usually simple, occasionally double or triple and very rarely fasciculate, with a cluster of 6 — 8 on a single stipe. Single sporangia globose or globose-turbinate ^ — § of a mm. in the transverse diameter. Colour of sporangia a dark nut-brown, which is uniform below, but checkered or broken above into irregular patches with broad septa of a bright yellow colour. Stipes about 1 mm. in height, rough or granular on the surface, quite thick and equal throughout their length, and dark brown in colour. Spores and capillitium concolourous, being a bright yellow colour in mass. Trklda. 185 Elatcrs cylindrical, 375 — 4 /x in diameter, terminating in sliort, sharp, smooth ends 4 — 6 /x long. Elater spirals 4 in number, spinulose with numerous irregular spines, coarse, wind- ing irregularly. Adjoining spirals often united with each other by interspiral branches which run either longitudinally or obliquely in the direction of the spirals. Spores 12 — 13 /x in diameter, delicately warted when examined under a high power lens. Shawangunk and Adirondack Mts., N. Y. This Trichia is conspicuous for the checkering or areolation of the upper surface in the perfectly mature sporangia, showing a sharp contrast between the adjoining nut brown and bright yellow colours. In this respect it resembles Tr. fi-agilis, which sometimes exhibits in its var. latcritia and also in one of its simple forms, a dull mottling of the colour of the upper siu-face of the sporangia. Trichia erecta, Eex, Proc. Acad. Sci. Phil., 1890, p. 193. Uxsicc.—EWk and Everh., N. Amer. Fung., Ser. II., n. 2496. Trichia Decaisneana, De Bary. Sporangia pyriform, brownish flesh-colour, shining, stipitate ; stem red, then blackish-brown, very much plicate, equal; mass of capillitium and spores yelloivish fiesh-colour ; inclosed in an inner mc/nhrane connate with the outer loall of the sporangium ; elaters cylindrical^ inflated near the tips and ending in smooth, tapering, curved spines 3 — 6 times as long as elater, spirals 5 — 6, fiexuous, spinulose^ in some cases parts of the elaters have the spirals in the form of distant ridges or wrinkles ; spores delicately warted, 10 — 11 /^ diameter. Trichia Decaisneana, De Bary, in Rost., Mon., p. 250, figs. 219, 220; Schroeter, p. 112; Cooke, Myx. Brit., figs. 219, 220; Mass., Mon. Trich., p. 338. On Jungermannia. Germany. According to Schroeter the spores measure from 13 — 14 fx, the elaters have 4 spirals, and are 4 — 5 mm. thick. As these measurements differ from those given by the author of the 186 A Monograph of the Myxogastres. species, the question that naturally suggests itself is, has Schroeter had the true species in view ? In Rostafinski's Monograph, fig. 220, the spirals of the elaters are represented as broad and fiat, no spines are shown in the figure. C. Spores with elongated, raised, fiat hmds not comhined to fm^iii a netioorh. § Bands not jntnctate. Trichia Balfourii, Mass. Sporangia sessile, base broad or narrowed, crowded, hemi- spherical or angular from mutual pressure, clear primrose-yellow ; mass of elaters and spores deeper and duller yellow; elaters cylindrical, 9 — 10 jx thick, sometimes swollen near the apex, which is abrwptly narrowed into from 1 — 3 sliort, smooth spines, generally more or less bent, spirals thin, not prominent, rather distant, furnished with scattered, rudimentary sp)ines ; spores globose, with a few broad, slightly raised, fiat lands, not punctate, nor combined in a reticulate manner ; 16 — 18 ju. diameter. Trichia Balfourii, Mass., Rev. Trich., p. 339, fig. 4. (Type in Herb., Kew.) On wood. Cape of Good Hope. Closely allied to T. JacJcii, but known by the absence of punctiform markings on the raised bands of the spores. In T. verrucosa the bands on the spores are much shorter and more numerous, the elaters of the last-named species also differ in the crowded spirals without spines and the simple stout tips. Trichia sulphurea, Mass. Sporangia densely crowded, subdepressed, circular, angular, or reniform in outline, pale yellow, smooth, mass of elaters and spores pale lemon-yellow; elaters cylindrical, simple, or fre- quently branched, especially near the tips, 9 — 10 /x thick, spirals crotvded, not very prominent, tips not thickened, smooth, acute, straight or slightly curved, 10 — 14 /u. long; spores globose, with numerous short, slightly raised, straight or crescent-shaped fi,at s, 10 — 14 iM diameter. Trichia. 187 Trichia sulphnrca, Mass., Rev. Trich., p. 330, fig. 3. (Type in Herb. Berk., Kew, n. 10906.) On wood. Ceylon. A very fine, large species with sporangia up to 1 mm. diameter, forming densely crowded patches of considerable size. Characterized by the thick cylindrical elaters with crowded spirals, and the numerous short flat bands on the spores, which look, like warts under a low power. Most nearly allied to T. nitens, but the spores of the latter have true rounded warts, and the spirals on the elaters are further apart. § Bands with minute dejjrcssions. Trichia abrupta, Cooke. Sporangia densely gregarious, sessile on a broad base, hypo- thallus firm, usually angular from mutual pressure, pale yellow ; mass of elaters and spores orange ; elaters cylindrical, 7 — 9 /x diameter, spirals rather distant, not prominent, with scattered, rudimentary spinules, tips usually smooth and equal in thick- ness to elater for a length of 8 — 10 ju., then terminating ahra^My in tioo or three tldn, straight or curved sjnncs 8 — 10 /x long; sjwres globose, toith numerous slightly raised, straight or c^irvcd short lands of irregidar width, furnished with miivute, scattered flits, 10 — 16 /x diameter. Trichia abrupta, Cooke, Ann. Lye. Nat Hist., N. York., vol. xi., No. 12, p. 404 ; Cooke, Myx. Brit., fig. 256 ; Mass., Mon. Trich., p. 16, f. 2; Sacc, Syll., vii., 1, n. 1511. (Type in Herb., Kew.) On wood. Britain (Ken Wood, Hampstead ; Kew, Carlisle) ; United States, Allied to Trichia Jachii, but distinguished by the more numerous and shorter bands on the spores and the different elaters; separated also from T. intermedia by tlie numerous hands on the spores not anastomosing, and by the absence of thin ridges parallel to the long axis of the elaters. 188 A Monograph of the Myxogastres. Trichia Jackii, Rost. Sporangia usually crowded, sessile on a narrow or broad base, liypothallus well-developed, circular, angular, or elliptical in shape, dull yellow ; mass of elaters and spores yellow ; elatcrs cylindrical, 5 — 7 ju, thick, tips smooth, acute, straight or a little bent, spirals not very prominent, distant, sometimes with rudimentary spinules ; spores globose, with scattered, slightly elevated, irregular, broad fiat hands, slightly curved or sinuous, not combined to form a network, surface of bands with minute, pits, 12 — 15 /x diameter. Trichia Jacldi, Rost., Mon., p. 258, f. 242 ; Cooke, Myx. Brit., f. 242; Schroeter, p. 113; Sacc, Syll., vii. 1, no. 1500; Raunk., Myx. Dan., p. 69, t. 4, f. 5 ; Mass., Mon. Trich., p. 16, f. 5. On wood and bark. Britain (Brighton, Highgate, Castle Howard, Yorks; Glamis, N. B.) ; Germany; Italy; Switzerland; Denmark. Allied to Trichia ahrupta, but distinguished by the fewer and longer bands on the epispore, and the undivided tips of the elaters. Trichia intermedia, Mass. (figs. 180 — 182). Sporangia subglobose, sessile on a broad base, crowded, often irregular from mutual pressure, smooth, shining, bright ochre ; mass of spores and capillitium clear pale chrome-yellow ; threads simple, cylindrical, 8 — 10 /x thick, ending rather abruptly in a short, smooth apiculus, spirals rather close, not prominent, sometimes branched, with a few rudimentary scattered spinules, and connected by thinner raised bands running more or less parallel to the long axis of the elater; spores globose, with a few distant raised flat flexuous bands that anastomose to form an irregular network, often leaving free ends, surface of hands ivith mimcte depressions usually arranged in a single row, 9 — 11 /ut diameter. Triehia intermedia, Mass., Rev. Trich., p. 341, pi. v., fig. 1. (Type in Herb., Kew.) On trunks. Scarborough, Epping Forest ; Finland. T rich id. 189 The spores somewhat resemble those of Triclna Jachii, Rost., but are smaller ; the raised bands are narrower and anastomose irregularly, and the minute depressions more constantly arranged in a single row. Agreeing with Trichia clirysospcrma, Rost, in having the spirals on the elaters connected by ridges, but at once distinguished by the bands on the spores being furnished with minute depressions. I was at one time led to believe, from examination of a specimen supposed to be authentic, communicated by Mr. H. Wingate, of Philadelphia, who received it from the author, that the present species was synonymous Avith Trichia 'p'^^oximcUa, Karst., but a second authentic speci- men of the last-named species agrees with the characters given by Karsten, hence T. intermedia will remain as a distinct species. Var. persimilis, elaters very much and irregularly branched, hranehes usually short, and as thick as the elaters; rest as in typical form. Trichia persimilis, Karst., in Not, Sallsk. pro Faun, et Flor. Fenn., Forh., 1868, ix., p. 353; Karst., Myx. Fenn., p. 139; Sacc, Syll., n. 1506 ; Mass., Rev. Trich., p. 338. On wood. Finland. An authentic specimen from the author proves the present form to be a variety, or more probably an abnormal form, of the present species. D. Spores loith raised fiat hands combined to form a network. * Bands -plain. Trichia chrysosperma, Rost. Sporangia crowded or scattered, sessile, base broad or naiTow, yellow or ochraceous cinnamon; mass of elaters and spores bright primrose-yellow; elaters cylindrical, 5 — 7 ix thick, tips short, smooth, straight or curved, spirals not very prominent, rather distant, sometimes with a few scattered spinules, con- nected ly thin ridges running parallel to the lang axis of the elater; spores globose, with deep, narroio raised hands combined to form an irregular polygonal netioork ; surface of lands not punctcde, 12 — 14 /x diameter. 190 A Monograph of the Myxogastres. Trichia chi-ysospenna, Rost., Mon., p. 255, figs. 209, 213, 240 Cooke, Myx. Brit., p. 65, figs. 209, 213, 240; Schroeter, p. 113 Sacc, Syll., vii., 1, n. 1498 ; Raunk., Myx. Dan., p. 69, t. 4, f. 1 Mass., Mon. Tricli., p. 17, f. 10. Exsicc. — Rab., Fung. Eur., n. 567. On wood, bai'k, moss, &c. Britain (Highgate) ; Germany. A rare species, characterized by the deep narrow ridges forming a network on the epispore and the surface of the bands not being punctate, and by the narrow, raised lines running parallel to the long axis of the elaters. It is amazing how Rostafinski managed to give so many synonyms for the present species, considering that the specific characters require for their elucidation at least a y^th oil immersion objective, yet this is done, and without any query as to their accuracy, although presumably not many of the types, if indeed any, were forthcoming. (Rostafinski's Synonyms.) Lycoperdon gregarium, Retz. Obs., i., 33 (1769). Zycoperdon favogineum, Batsch., f. 173 (1786). Stcmonitis i^yriformis, Roth. Fl. Germ., i., 548 (1788), SphaerocarpiLs chrysospcrinus, Bull., t. 417, f 4 (1791). Stemonitis favoginea, Gmel. Sys., 1470 (1791). Trichia nitens, Pers. Obs., i., 62 (1796). Trichia favoginea, Pers. Disp., 10 (1797). Trichia chrysosperma, DC. Fl. Fr., 673 (1805); Eng. Fl., v., 320; Cooke, Hdbk., No. 1187; Fung. Britt., ii., 524, 527. Trichia turhinata, Purt, Brit., ii., 1115 (1817). Claihroides fluvesccns, Hall. t. ], f 7 (1742). Trichia, Hall, 2168, t. 48, f. 7 (1768). Lycopcrclon aggrcgatum, Retz, Fl. Scan., 1627 (1769), Lycopcrdon cpiphyllam, Light. Fl. Sc, 1069 (1777), Glathrus turhinatus, Huds. Fl. Ang., 632 (1778) ; Bolt., t. 94, f. 3. Trichia pyriformis, Vill. Fl. Dauph., 1060 (1789). Stemonitis pyrifm-mis, Pers., in Gmel. Sys., 1468 (1791). Trkhia. 191 TricJiia tiirUnata, With. Arr., iv., 480 (1702) ; Sow., t. 85 ; Eng. FI., v., 320; Cooke, Hdbk., 118G. Trichia jvjrifoo'mis, Pers. Disp., 10 (1797). Trkhia olivacca, Pers. Obs., i., 62 (1796), in part. Trichia ovata, Pers. Obs., ii., 35 (1796) ; Schum. Saell., 1454 ; FI. Dan., t. 1313, f. 1. Trichia vulgaris, Pers. Obs., ii., 32 (1799). rhysarum contextum, Spr. Sys., ix., 20 (1817). Trichia verrucosa, Berk. Sporangia fyriform, brown or chestnut, shining, passing down- wards into a long, slender stem, simple or botryoid, scattered, springing from a thick, Iroadly effused hypothallus ; mass of capillitium and spores ochraceous ; threads of capillitium not branched, cijlindrical, 8 — 10 ii thick, with smooth, tapering tips of variable length, and straight or curved, spirals close, thin, not prominent ; spores globose, with 7iarroiv, raised fiat hands comhined into a few large, irregular, polygonal meshes, hands not 2mnctate, 14 — 16 ix diameter. Trichia verrucosa. Berk,, Flor. Tasm., p. 269 ; Mass., Rev. Trich., p. 343, fig. 9. (Type in Herb. Berk., Kew, n. 10906.) On wood. Tasmania. Differs from T. chrysosperma and T. dictyospora in the scat- tered sporangia springing from a stout hypothallus, and also in the characters of the elaters and spores. From 2 — 3 mm. high. Usually not more than one complete polygon is present on a hemisphere of the spore. Trichia Kalbreyeri, Mass. Sporangia crowded, sessile, often irregular from mutual pressure, pale yellow, smooth ; mass of capillitium and spores pale primrose-yellow; threads of capillitium cylindrical, 9 — lOjix thick, with short, smooth, tapering ends, spirals not 2Ji'ominent, thin, close ; spores globose, with raised, narroiv fiat hands form- ing an irregular, polygonal network, bands not punctate, 11 — 14 /x diameter. 192 A Monograph of the Myxogastres. Trichia Kcdhreyeri, Mass., Rev. Trich., p. 344, fig. 8. On wood and living leaves. New Granada. Externally resembling T. ckrysospcrma, but known by the absence of ridges between tlie spirals, and the bands forming more numerous polygons, 2 — 3 complete ones being present on a hemisphere of the spore. Trichia scabra, Kost. Sporangia gregarious or scattered, attached by a broad base to a hypothallus, circular or irregular from mutual pressure, varying from pale yellow, through dirty orange to brown ; mass of elaters and spores clear orange ; elaters cylindrical, 6 — 8 jx thick, ending in smooth, acute, straight or slightly bent tips 7 — 10 IX long, spirals not very prominent, rather distant, hearing numerous short, acute, straight spines; spores globose, covered with a very fine networlz of raised lines, 8 — 12 \i. diameter. Trichia scalra. Host., Mon., p. 258, figs. 214—217, 239; Cooke, Myx. Brit, figs. 214—217, 239; Mass., Mon. Trich., p. 13, fig. 13 (spore incorrectly drawn with a warted instead of a very delicately reticulated epispore) ; Schroeter, p. 113; Sacc. Syll., vii., n. 1500 ; Eaunk., Myx. Dan., p. 68, t. 4, f. 2. Trichia scahra, v. aurea, Cke., Myx., U. States, Ann. Lye. Nat. Hist., N. York., v. xi.. No. 12, p. 403. Exsicc. — Roum., Fung. Gall., 1005; Ellis and Everh., N. Amer. Fung., 2100. On wood, moss, &c. Britain (Queen's Cottage Grounds, Kew, Birmingham ; Taunton, Notts ; Scarboro' ; Carlisle) ; France ; Germany ; Denmark ; U. States ; Ceylon ; Australia. Var. analogia. Spores on spirals of elaters rudimentary or absent. Cke., Myx., U. States, Ann. Lye. Nat. Hist., N. York, vol. xi.. No. 12. p. 403. Britain (Carlisle) ; U. States. Trichia fallax, Rost. Sporangia pyriform, stipitate or subsessile, ochraceous, some- times tinged olive, dull or shining; stem dark, usually longi- T rich id. 193 tiulinally wrinkled, filled with large, globose, suhcmgalar cells wliieh become smaller icjnvards and pass into normal spores; elaters pyriform, simple or branched, 5 — 6 fx thick, ending in long, smooth, tapering tips, spirals rather close, thin, not pro- minent; spores globose, epispore, covered with a very fine, irregular network, 10 — 12 n, diameter. Trichia fallax, Rost., Mon., p. 243, figs. 211, 221, 222, 233— 236; Cke., Myx. Brit., p. 61, figs. 211, 222, 233—236; Sacc, Syll., vii., 1, n. 1493 ; Mass., Mon. Trich., p. 8, figs. 21 and 27 (the spores incorrectly represented as warted instead of being delicately reticulated); Schroeter, p. Ill; Raunk., Myx. Dan., p. m, t. 4, f. 4. Exsicc. — Fuckel, Fung. Rhen., 1435 ; Jack, Leiner u. Sitz., 420; Rab., Fung. Eur., 1666; Mong. and Nest., 284; Roum., Fung. Sel. Gall, 42. On rotten wood. Britain (Bristol, Kew, King's Cliffe, Norths ; Carlisle, Scarboro', Linlithgow, Glamis, N. B.) ; France ; Germany ; Switzerland ; Denmark ; United States ; Cuba ; Venezuela. Distinctly marked amongst the species with fusiform elaters by the stem being filled with large cells, and the very delicately and minutely reticulated spores. (Rostafinski's Synonyms.) Mucor capitulis pyriformis, Fl. Dan., t. 647, f. 2 (1770). Mucor miniatus, Jacq. Misc., t. 290 (1778). Stemonitis fiavcscens, Schrank., p. 19 (1792). Lycoperdon aggregatum, Liljeb. Fl. Scan., 460 (1792). Lycoperdon p)usilhim, Hedw. Abh., t. 3, f. 2 (1793). Trichia fallax, Pers. Obs., iii., t. 4, 5 (1797); Nees, f. 113; Corda Ic, iv., 97; Eng. Fl., v., 319; Cooke, Hdbk., 1182. Fhysamm pyriforme, Schum. Saell., 1448 (1803). Trichia virescens, Schum. Saell., 1459 (1803). Trichia cerina, Ditm., t. 25 (1817); Curr. Micr. Journ., v., p. 127; Cooke, Hdbk., No. 1184. Trichia ficlva, Purt. Mid. Fl., 1534 (1817). Trichia clavata, Wigaud, No. 3 (1S63). 194 A Monograph of the Myxogastres. TricUa furcata, Wigand, No. 4 (1863). Arcyria clongata, Bong. Herb. * Bands icith minute depressions on the surface. Trichia afiinis, De Bary. Sporangia clustered, circular or elliptical, sessile on a broad base, seated on a distinct hypothallus, clear pale yellow ; mass of elaters and spores pale yellow; elaters cylindrical, 4 — 6 ii thick, ending in short, tapering, smooth tips, spirals thin, rather close, not prominent, sometimes with short spinules ; spores globose, with Iroad, slightly raised fiat lands comhined into a network, or sometimes ivith free ends, surface of hands ^9t^li of the Myxogastres. threads often forked at acute angles, anastomosing laterally, hrotviiish-lilac, furnished here and there with thickenings ; spores dusky violet, globose, with 'j)rominent, elongated, Jicxuoiis ridges, 8 — 11 /x diameter. " Didymium dealhatum, B. and C," in Herb. Berk., n. 10756. Chondrioderma siibdictyosijermuin, Rost., Mon. Append., p. 16; Sacc, Syll., n. 1260. On wood, amongst moss. Venezuela. A very fine and distinct species, sporangium pure white, polished, about "5 mm. diameter. There appears to be no good reason why Berkeley's specific name, although only a manu- script one, should have been changed by Rostafinski. The description of the present species in Saccardo's Sylloge is meaningless. (Rostafinski's Synonym.) Didymium dealhatum, B. msc. Chondrioderma Friesianum, Rost. Sporangia sessile, subhemispherical, wall very much charged with lime, snow-white, after the crust has fallen away, grey ; columella distinct, lenticularly depressed, yellowish-gilvous or flesh-colour ; capillitium well-developed, colourless, threads com- bined to form a network ; spores pale violet, 8 //, diameter, smooth. Chondrioderma Friesianum, in Fuck el's Sym. Myc, 2, p. 74<; Rost., Mon., p. 172 ; Sacc, Syll, no. 1266. On bark, stems, &c. Europe. Distinguished from Chondrioderma glohosum by its irregular form, by the inside of the sporangial wall subreticulate, and the almost umber spores. (Rostafinski's Synonyms.) Diderma difforme, Sommf , Fl. Lap., p. 217 ; non Pers ; sed Fr., S. M., iii., p. 106 (1825). Chondrioderma Friesianum, Rost., in Fuckcl's Symb. Myc, 2, Nach., p. 74. CliondriodeniKi. 209 Chondrioderma albescens, Phil. Sporangia fjlohose or slightly depressed, sessile on a hrond hasc^ outer wall crustaceous, white, polished, becoming irregularly ruptured above, inner wall very thin, ochraccous towards base of sporangium and where it passes over the columella, colour- less and very delicate above; columella well-developed, more or less globose, ochraccous ; mass of spores, blackish-purple ; 4;hreads of capillitium fuliginous or almost colourless, about 3 /x thick at the base, remaining for some distance simple, then branching at acute angles, towards the apices anastomosing freely to form a dense, irregular net; spores globose, lilac-brown, minutely vcrrucidose, 10 — 13 /x diameter. Chondrioderma albescens, Phil., in Herb. Diderma albescens, Phil., Grev., v., 5, p. 114, t. 87, f. 3, a — f ; Sacc, Syll, 1291. (Type in Herb. Phillips.) On pine bark. California. Gregarious, or towards the margin of the clusters, scattered, 1 — 1*5 mm. diameter. After dehiscence, when the spores are blown away, the bright ochraceous columella and base of sjiorangium are very conspicuous. The threads of the capil- litium are sometimes furnished with scattered swollen portions. Chondrioderma simulans. Rest. Sporangia spherical, narrowed at the base, adnate to a well- developed, chalk-white hypothallus composed mostly of lime, outer wall crustaceous, clialk-ichitc, inner wall cinereous or variously bright-coloured ; columella generally very small, chalk- wliite, spherical or conical ; threads of capillitium forming a dense net, with triangular protuberances at the nodes, brownish- violet ; spores dingy violet, very spinulosc, 1 2'5 ix diameter. Chondrioderma simulans, Rost., Mon. Append., p. 20 ; Sacc, Syll., 1270. Chondrioderma Saundersii, B. and Br. Sporangia scattered, sessile on a broad base, \vy\ much 210 A Monograpk of the Mijjcogastres. depressed, circular in outline, wall veiy thin, chalk-white, polished, breaking away in irregular patches ; cohimcUd flattened, v:hitc, small, mass of spores blackish-puiiDle ; threads of capil- litium equal, about 2 ju, thick, frequently branching and com- bined laterally to form a dense net; spores globose, smooth, dingy violet, 9 — 10 fx diameter. Chondrioderma Saundersii, B. and Br., in Herb. (Type in Herb. Berk., no. 10744.) On living fronds of ferns. Java. Resembling superficially circular forms of Chondriodcrmct dif- formc,h\xi distinguished by the columella and dense capillitium. Sporangia 2 — 3 mm. diameter, outline circular, often waved; a very thin powdery layer of minute granules of lime is frequently present on the mass of spores, which then appears dark grey. Chondrioderma afRne, Rost. Sporangia small, densely crowded on a common hypothallus, irregularly angular, depressed, surface smooth, greyish-white; columella irregular, small, white, rarely very conspicuous; threads of capillitium thin, forming a net, dirty-violet, often furnished with fusiform subviolaceous protuberances; spores subviolaceous, densely aculeate, 10 — 14 /x diameter. Clwndriodcrma ajfine, Rost., Mon. App., p. 18; Sacc, Syll,, 1271. Poland ; Germany ; France. Appears to be very closely allied to Didymium sjmmarioides of the present work (Chondrioderma spumarioides, Rost.) ; differ- ing in the smooth surface of the sporangial wall, the dingy colour of the capillitium, and the lai-ger, densely aculeate spores. * * Sjwranfjiiim coloitrcd. Chondrioderma testaceum, Rost. Sporangia gregarious or crowded, sessile on a broad base, hemispherical or elliptical, rather depressed, sometimes irregular from mutual pressure, outer wall smooth, from hrich-rcd to almost ChondvioiJcnna . 211 colunrh-Hii, dehiscing irregularly, inner wall violet- r/ri\i/ ; threads of capillitium slender, brownish-lilac or almost colourless, fre- quently bifurcating and anastomosing, sometimes flcxuous; columella sibhglohose or dcjn'esscd, reddish or almost colourless; rugose, spores globose, brownish-lilac, minutely vcrruculose, 9 — 11 fi diameter. Ghondrioilcrma testaceum, Host., Mon., p. 179, figs. 135 — 136 ; Schroeter, p. 135; Sacc, Syll., n. 1274; Cooke, Myx. Brit., figs. 135, 136. JExsicc. — Fuckel, Fung. Rhen., 14G7 ; Dcsm., Cr. Fr., Scr. I., 706 (as Diderma testaceum) ; Ellis and Everh., N. Amer. Fung., 2093. On mosses, leaves, twigs, &c. Sweden ; Germany ; Finland ; Hungary; United States. Varying from 'o — 1 mm. diameter; the colour of the spor- angium bright brick-red, but it is sometimes much paler. (Rostafinski's Synonyms.) Bcticularia spJiaeroidalis, Var. 2, Bull. Champ., p. 94, t. 446, f. 2, D (1791). Didymium testaceiim, Schrad., p. 25, t. 5, f. 1, 2 (1897). Diderma testaceum, Pers. Syn., p. 167 (1801). Gionium testaceum, Spr. Syst., iv., p. 529. Chondrioderma sublateritium, Rost. Densely gregarious, sessile on a broad base, but rarely deformed by crowding, circular in outline, depressed, umlilieate ahote, X)cde Irich-red, jyolishcd ; inner wall lead-colour ; columella small, Jlattened, reddish, capillitium forming a flaccid network, threads thin, sometimes flexuous, with here and there small fusiform swellings without lime ; spores globose, pale lilac-brown, minutely vcrruculose, 9 — 11 fx diameter. Chondrioderma suhlateritimi, B. and Br., Journ. Linn. Soc, vol. xiv., p. 82. (Type in Herb. Berk., n. 10728.) On leaves. Ceylon. 212 A Monogrcrpfi of the Myxogastres. Closely allied to Chomlrioderma tcstaccum, if indeed it is more than a geographical variety. Sporangia flattened, umbilicate above, frequently dehiscing in a circumscissile manner, Chondrioderma mutabile, Schroeter. Sporangia sessile, deformed, hemis2')hcrico-dciirc&sed, or elongated and curved, reniform, semicircular, &c., 1 — 3 /x long, 1 mm. broad, hard, fragile, bright, greyish-hroivn ; columella tvcll de- veloped, same shape as the sporangium, bright rufaiis-hroum ; threads of capitulum thin, violet., here and there thickened in a nodulose manner; spores globose, 11 — 14 \x diameter, blackish- violet, mucronulate. Chondrioderma mutaUle, Schroeter, Kr. Fl., p. 123; Sacc, Syll., 1273. On rotten wood. Silesia. Chondrioderma fallax, Rest. Sporangia spherical, sessile, yellowish-vMte, seated in numbers, but not crowded, on a distinct common hyjriothallus ; columella either minute and flcdtened,or of medium size and ovate; threads fasciculate for some distance then branching copiously, obscure violet; spores obscure violet : ivaoied, 12 — 14 /x diameter. Chondrioderma fallax, Rest., Mon., p. 171 ; Sacc, Syll., no. 1261. Salzburg, Tyrol. B. Columella ahsent. Chondrioderma difForme, Rost. (figs. 35 — 38). Sporangia sessile on a Iroad base, convex, circular or irregularly elongated, outer wall snow-white, rather thick, breaking away in patches, inner wall separated from the outer by a space containing air, thin, without lime, brownish-ochre, sometimes iridescent ; mass of spores blackish ; columella absent or repre- sented by a small accumulation of lime at base of sporangium, which is covered by the inner wall ; capillitium scanty, some- times almost o\i?>o\Q,iQ,thrccuh springing from base of sporangium, Chondriodennd . 213 3—4- ^ at base, sliyhtly attenuated upwards, forked, pale brown or colourless; spores globose, smooth, dingy violet, 10 — 13 /x diameter. Ghondriodcrvia difforme, Rost,, Mon., p. 177, fio-s. 137, 164, 165; Cooke, Myx. Brit., p. 39, figs. 137, 164, 165; Sacc.,' Syll.,' 1282; Raunk., 83. CJwndrioderma Cnhcnsc (B. and C), Rost., Mon. Append., p. 19; IHderma Cuhcnsc, B. and C, Journ. Linn Soc v x p. 347. ' ' " Ex8icc.—Uh., PI. Crypt. Ard., Fasc. III., 276 (as Didervia liceoides)', Fckl, Fung. Rhen., 2300 (as Didcrma liceoidcs)- Rabh., F. Eur., 1423 (as Didymiitm Lihertianum, De Bary); Rab.-Wint., Fung. Eur., 3172; Roum., Fung. Gall., 1311 and 2956; Rab., Herb. Myc, 456; Roum., Fung. Gall., 243 (as Diderma candidum); Sydow, Myc. March., 1497; Desra., Cr. Fr., Ser. I., 370 (as Diderma difforme); Fuckel, Fung. Rhen., 1464 (as Leocarpus calcareus, Link); Ellis, N. Amer. Fung., 1217; not typical, the spores smaller than in type, and the inner wall bright grey. On twigs, leaves, &c. Britain (Elmsted, Kew, Northampton, Rudloe, Twycross, Carlisle, Scarboro', Linlithgow, Appin, N. B.) ; France ; Germany ; Switzerland ; U. States ; India ; Australia. Sporangia 1 — 2 mm. across when circular, hemispherical in section. (Rostafinski's Synonyms.) Fu'ticularia angulata, Pers. in Gmel, p. 1472 (1791). Didcrma difforme, Pers. Disp., p. 9 (1797); Icon. Pict., t. 12 f. 3— 5; Nees, f. 105. Lieca caesia, Schum. Saell, 1500 (1803). Physancm difforme, Link, Diss., i., 27 (1809). Am2jhis!porinm versieolor, Fr., Gast., 19 (1818). Licea alba, Nees, in Kze., Myk., Heft., ii., 66 (1823). Lyeorjala mimdum, Grev., S. C. Fl., t. 40 (1823). Betieularia imsilla, Fr., Orb. Vet., i., 147 (1825). Diderma eyanescens, Fr., S. M., iii., 109 (1829); Eng. Fl., v., p. 312; Cooke, Hdbk., No. 1115. 214 A Monograph of the Myxogastres. Physat'um caesium, Fr,, S. M., iii., 147 (1829). Physaimm alhnm, Fr., S. M., iii., 147 (1820); LetelL, t. 710, f. 4; Cooke, Hdbk., No. 1140. Didymium difforme, Duby, Bot. Gall., ii., 858 (1830). Didcrma nitcns, Klotzsch, in Hook. Herb.; Eng. FL, v., p. 312; Cooke, Hdbk., 1113. Didcrma Ncesii, Corda, Ic, ii., f. 58 (1838). Leocarpus cyanesccns, Fr., S. V. S., 450 (1849). Zeocarpics mtem, Fr., S. V. S., 450 (1849). Didcrma Lihcrtianum, Fres., Beit., t. iv., f. 16 — 27 (1850). Didymium Lihcrtianum, De Bary, Mycetozoa (1864). Chondrioderma Berkeleyanum, Rost. Sporangia slightly scutcllate, snow-white, stipitate ; stem rigid, hay at the hase, hecoming jJaler upivards; threads of capillitium slender, anastomosing to form a net ; spores dingy violet, strongly vxcrtcd, 10 — 11 II diameter. Chondrioderma Bcrkelcyanum, Rost., Mon. App., p. 16 ; Sacc, Syll., no. 1258. Island of Tahiti. There is no specimen in the Berkeleyan Herbarium answer- ing to the above description, and in the " Introduction to Cryptogamic Botany," p. 335, the spores are described and figured with a reticulate epispore. (Rostafinski's Synonym.) Tricha7iiphora ]iczizoidcs, Berk., Intr. Cr. Bot., p. 335, non Jungh. Chondrioderma physarioides, Rost. (figs. 59 — 62). Gregarious, sessile on a broad base, circular or irregularly elongated, depressed, outer cretaceous wall polished, pure white or ivory tinted, inner wall dirty ochraccons, sometimes separated from the outer wall by a space containing air ; columella ohsoletc ; threads of capillitium equal, about 3 jw thick, colourless, some- times rough in parts with minute particles of lime, hranchi^ig Chondriodcrmd. 215 ioTefj/nlarli/ and foi-ming a flaccid net; spores globose, dingy violet, minuteljj verruculose, 12 — 13 [j. diameter. Chondriodernia 2^hysarioidcs, Rost, Mod., p. 170; Cooke Myx Brit., p. 37. Cho')id,Tiodemia dtylanatum, Rost., Mon. Append p 17 • Sacc Syll.. 1264. ' ' ' On rotten wood, leaves, &c. Britain (King's Cliffe, Black- heath, Carlisle) ; France ; Switzerland. Sporangia 1 — 3 mm. when circular, from 8—5 mm. when elongated ; in the elongated form the sporangium is frequently folded on itself, and sometimes it takes two or three spiral turns, leaving a minute cavity in the centre. Distinguished from Chondriodcrma difforme by the warted spores and large size of the sporangia, and from 0. nivetim by the absence of tlie columella. Sometimes the whole of the outer calcareous coat breaks away in a circumscissile manner close to the adnate Chondriodcrma crustaceum, Berl. Effused or circumambient, crowded, sessile, subglobose, smooth, white, outer peridium crustaceous, like the shell of some small egg, the inner delicate, appearing cinerous to the naked eye, iridescent under the microscope; columella none; spores globose, black, -0005 in. in diameter ( ^ about 13 ju). Chondriodcrma crustaceum, Berlose, in Sacc, Syll., no. 1290. Didcrma crustaceum, Peck, in 26th Report of State Mus. N. Y., p. 74. On fallen sticks and leaves. Memphis ; United States. Owing to absence of information respecting the capillitium and presence or absence of markings on the epispore, the p(jsitiun of the species is uncertain. Chondrioderma liceoides, Rost. Plasmodium scarcely as thicli as a needle, creepmff or x>ulvinate, wall highly charged with lime, wliite externcdly, inside reddish- brown ; inner wall contiguous to outer, delicate, viokt ; capil- litium f.iirly abundant, threads swq^le, or once or ticiec diehoto- 216 A Monograjyh of the Myxogastres. maics near the ajKx, spores smooth, obscure violet, 11 — 12 /ut diameter. Chondnoderma liccoides, Rost, Mon. Append., p. 17; Sacc, Syll, 1262. On rotten jiourd. Carolina. (Rostafinski's Synonyms.) Licca macrosperiiia, Sz., Am. Fung., no. 2317 (1834). Ligynota nigra, Fr., Sra. Veg. Scan., p. 459 in notam. (1840). Chondrioderma reticulatum, Rost. Plasmodium sessile, vein-like, flattened, arcuate, comhined into an irregular nctworic, seated on a whitish, delicate, reticulated, S2')reading hypothallus ; columella absent ; threads of capillitium colourless, veo^y thin, comhined to form a dense net ; sjjores smooth, obscure violet, 7 — 8 /x diameter, Chondrioderma reticulatum, Rost., Mon., p. 170. On rotten branches of pine. Switzerland. I have not seen the present species, but the description seems to suggest that it may possibly be a plasmodiocarp form of C. Miclielii. * * S'porangiitm coloured. Chondrioderma ochraceuin, Schroeter. Sporangia sessile, irregular, globose, reniform, semicircular, &c., 1 — 2 mm. long, 1 mm. across, densely gregarioics, outer wall oehraceous-brown, falling away in irregular squamules; inner wall thin, colourless; columella cd)sent ; capillitium copious, threads- 2 — 3 fx thick, violet, smooth, straight or flexuous, frequently anastomosing, and here and there forming a dense reticulation; spores globose, indistinctly i)unctulatc, blackish- violet, 9 — 11 ix diameter. Chondrioderma ochraceum, Schroeter, Kr. FL, 124 ; Sacc, Syll., v., 7, part, I., no. 1283. On hepatics. Silesia. Chond node rum. 217 Chondrioderma Sauteri, Rost. Sporangia sessile, irregularly elli'iisoid, dejyorsscd, Uood-red, shiniui/ ; iDiier wall wliitisli-brown, dull; columella alsent, threads of the capillitium fasciculate but not forming a net; spores pale violet, verruculosc, 8 — 12 [x diameter. Chondrioderma Sauteri, Host., Mon., p. 181 ; Sacc, Syll., 1281. Tyrol. There is some difficulty in ascertaining the exact meaning of the colour of the sporangium as given by Rostafinski, who calls it " kawowo-mleczne." It is translated in Saccardo's Sylloge as " sanguineo-lacteis." Chondrioderma vaccinum, Rost. Sessile, adnate, orbicular, obiter wall thick, adnatc, inner very thin, evanescent ; columella ahsent ; threads of capillitium hyaline, delicate; spores pale brownish-violet, 10 — 11 ij. diameter, minutely wavted. Chondrioderma vaccinum, Rost., Mon., p. 180 ; Sacc, SylL, no. 1275. Diderma vaccinum, Dur. et Mont., Fl. Alg., p. 407, t. 22 bis., f. 1 ; Mont, SylL, n. 1071. (Type in Herb. Mus., Paris.) On fallen brandies of Ojyuntia. Algeria. Externally bearing a close resemblance to Chondrioderma testaeeum, but easily recognized by the absence of a columella; Chondrioderma sMateritiwm is separated from the present species in having the upper surface of the sporangium broadly um- bilicate, and in the presence of a columella. Chondrioderma simplex, Schroeter. Sporangia sphaeroid, slightly flattened, scattered, simple, rigid, fragile, bright chestnut ; columella ahsent ; tubes of capil- litium radiating, repeatedly dichotomous, violet; sjyores smooth, bright violet, 7 — 9 fx diameter. Schroeter, Kr. Fl. Schles. Filze, p. 123; Sacc, Syll., 1272. On dead trunks. Silesia. 218 A Monoprffph of tJie Myxogastres. Chondrioderma calcareum, Rost. SporaDgia sessile and variahlc in form, ovate, elongated and curved, &c., convexo-depressed, glaucous and hyaline when moist, ajiaque and reddish or ^mZc v^hen dry, very fragile, at length rupturing irregularly, leaving a brow^n impression of the base after falling away; spores 9 — 11 ju, diameter, mixed with a few black threads. Chondrioderma calcareum, Host., in Fckl. S. Myc, 2 Nch., p. 74; Rost, Mon., p. 179; Sacc, SylL, 1280 ; Schroeter, p. 124. On dry stems. Germany. (Rostafinski's Synonyms.) Zcocarjms calcareus, Link, Diss. I., ]. c, p. 23 (1809). Diderma liceoides, Fr., 1. c, iii., p. 107 (1829). Diderma chalyheum, Wein., 1. c, p. 592 (1836). Diderm.a deplanahtm, Fckl, S. M., p. 341 ; Non. Fr. (1869). Chondrioderma calcareum, (Lk.) in Fckl., S. M., ii., Nch., p. 74 (1873). Chondrioderma Stahlii, Rost. Sporangia spherical, slightly flattened below, either dull whitish-brown or shining and dingy brown; dehiscing by a central pore, a long slit, or irregularly; stem hroivn, shining; columella absent ; threads of capillitium either simple or more or less repeatedly bifurcate above, net cmiihining to form a network; spores pale violet, minutely verruculosc, 9 /^ diameter. Chondrioderma Stahlii, Rost., Mon., p. 185, f. 168; Cooke, Brit. Myx., fig. 168; Sacc, Syll., 1287. Germany. DiDYMiUM, Schrad. (in part). Sporangia stipitate or sessile, wall single or double, surface covered with lime, either in the form of a pulverulent stratum of amorphous particles or crystals, or compacted into a granular (but not porcelain-like) crust; columella present or absent; Didymium, 2 1 9 capillitiunr usually well-devoloped, threads thin, without lime, coloured or colourless, either radiating from the columella or base of sporangium as simple or furcate threads, which usually combine laterally towards the apex, or combine throughout their length to form a dense, irregular network, with the anodes more or less triangular and flattened. Spores lilac or violet- brown. Didymium, Schrad., Nov. pi. gen., p. 20. Rost., Mon., p. IGO (in part); Cooke, Myx. Brit., p. 30 (in part); Sacc, Syll., v., 7, pt. I, p. 376 (in part); Zopf, p. 150. The leading idea of the genus as defined above is, the wall of the sporangium with lime outside, the thin threads of the capillitium not containing lime, either radiating, subsimple or bifurcating, or anastomosing irregularly throughout their length and forming a net as in the subgenus Hcmididymiuiii, the species of which are distinguished from those of Tilmadoche by the absence of well-defined, small, usually fusiform knots containing granules of lime. Thickenings are not uncommon in the capillitium of various species of Didymium, but lime is never present. Distrih. Temperate and tropical regions. Species 50. Sub-Gen. Didymium. Threads of capillitium almost simple, or bifurcating at acute angles, here and there connected laterally, but not forming a uniform net throughout the capillitium. A. S'porangium stipitate. (Sessile forms occur in D. squamulomm and D. farinaccum.) * Spores wartcd. Didymium farinaceum, Schrad. (figs. 29—35). Sporangium hemispherical, usually a little depressed, more or less umbilicate below, at first white with a continuous crust of lime, which soon becomes broken up into white ghstening granules scattered on the inner dark, wrinkled wuU ; stem 220 A MonograpJi of the Myiognstres. variable in length, blackish or rnfmis, wrinkled longitudinally, equal or slightly attenuated upwards, expanding at the base into a rudimentary hypothallus; coluinella hemispherical or depressed, covered with the inner darlc hrmun wrinkled skin, lacunose, the cavities filled with amorphous masses of lime; capillitium copious, threads springing from the columella, 2 — 3 /x thick, almost equal, dividing in a dichotomous manner, and becoming attached to the inner wall of the sporangium, the branches often connected laterally, and often more or less studded with very minute granules of lime, varying from colour- less, through pale brown lo dingy violet ; spores globose, dirty brownish-purple, minutely warted, 1 — 1 3 ju, diameter. a. gemoinum. Stem blackish or brown, as long or longer than the height of the sporangium. /3. suhsessile. Stem short, almost or altogether concealed in the umbilicus of the sporangium. Sporangia solitary or more or less confluent. y. nigrum. An abnormal form without lime, hence the sporangium from the first blackish. Stem short. Didymium farinaccum, Rost., Mon., p. 154, figs, 128, 171, 174; Cooke, Myx. Brit., p. 31, figs. 128, 171, 174; Sacc, Syll., 1039 ; Schroeter, p. 121 ; Raunk., Myx. Dan., p. 89. Didymium lohatum, Ft. (specimen named by Fries, in Herb. Berk., n. 10753). Exdcc. — Jack, Leiner u. Sitzenb., 424 ; Cooke, Fung. Brit., Ed. II., 521; Fuckel, Fung. Khen., 1461; Klotzsch., (Rab.) Herb. Myc, 138; Rab., Fung. Eur., 369. On wood, bark, leaves, living moss, &c. Britain (Chislehurst, Kew, Carlisle, Scarboro', Durham, Lin- lithgow, Appin) ; Sweden ; Germany ; Bavaria ; France ; United States ; S. W. Australia ; Benin Islands. Sporangia b — 1 mm. across, sometimes lobed, a character which gave origin to the spurious species D. lohatum; some- times two or several sporangia coalesce, the stems remaining distinct. The stem varies considerably in length, even in the same group, thus showing the small amount of value to be attached to Rostafinski's forms; in some specimens the stem Didymiam. 221 is three times as long as the height of the sporangium, in others so short as not to project beyond the umbilicus or depression at the base of the sporangium. The threads of the capillitium are often flexuous. The cokimella is at first dusky owing to being covered with the dark inner skin of the spor- angium, but in old plants this pellicle frequently disappears, leaving a white mass of lime, hence the columella appears white. (Rostafinski's Synonyms.) S'phacroccplmlus nigcr. Hall, t. 1, f. 2 (1742). Trkhia, Hall, No. 2160, t. 48, f. 2 (1768). M'ucor si^hacrocephcdus, Batsch., p. 157 (1783). Clathrus S2Jhaerocc2)hahcs, .'Rehl. (1786). Trichia glohosa, Vill., Fl. Dauph., 1061 (1789). Beticularia Jicmisj^herica, Bull., t. 446, f. 1 (1791). Trichia cojnjrressa, Trent, p. 229 (1797). Trichia sj)hacrica, Trent, p. 230 (1797). Trichia dcpressa, Trent, p. 231 (1797). Physarum mclanosjyervuwi, Pers. Disp., p. 8 (1797). Lidymium farinacmm, Schrad., t. 3, f. 6 (1797) ; Engl. Flor., v., 313; Cooke, Hdbk., No. 1123; Fung. Brit., Ed. H., 521.' Trichia sphacrocei^hala, Sow., t. 240 (1799). Trichia farinacca, Poir., EncycL, viii., 53. Physarum farinacmtm, Pers. Syn., 174 (1801). Physarum cinerasccns, Schum. Saell., 1426 (1803). Physarum dextrcssttm, Schum. Saell., 1439 (1803). Physarum glohosiwi, -Schum. Saell., 1442 (1803). Physarum oxyacanthae, Schum. Saell., 1427 (1803). Physarum cinercum, multis ? Physarum clavus, Link, Diss., i., 27 (1809). Physarum sinuosum, Link, Diss., i., 27 (1809), Physarum capikchirn, Link, Diss., i., 27 (1809). Diderma muscicola., Link, Diss., i., 27 (1809). Bidymiuiii ca2ntcdum, Link, Diss., iii., 27 (1816). Didymium lokdiim, Nces, f. 104 (1817); Cooke, Hdbk No 1129. 222 A MonoyrapJi of the Myocogastres. Didymium physarioides, Klotzsch. Strongylium minor, Fr., Gast., p. 9 (1817). Physarum melanopus, Fr., Gast., p. 23 (1817). Conium lobahtm, Spr., Syst., iv., 529 (1827). Didymmm marginatum, Fr., S. M., iii., 116 (1829). Didymium mdanopus, Fr., S. M., iii., 114 (1829); Berk., Ann. N. H., No. 882; Cooke, Hdbk., No. 1118. Didymium licmisphcricum, Fr., S. M., iii,, 115 (1829). Physarum nigrum, Fr., S. M., iii., 146 (1829); B. and Br., N. H., No. 1598; Grevillea, v., p. 12. Cionium farinaceum. Link, Hdbk., iii., 416 (1833). Didymium filamentosum, Wallr., 2187 (1833). Didymium affine, Raunk. Sporangia spherical-hemispherical, stipitate. Stetn thin, of equal length or longer than the sporangium, expanded into a circular hypothallus at the base, bright brown. Wall grey, withrmt lime, under the microscope colourless. Columella glo- bose-semiglobose, with the colour of the stem or brighter. Threads of the capillitium nearly hyaline, expanded into numer- ous shortly fusiform, Iro'ivnish-violet sioellings. Spores smooth or delicately warted, 8 — 9 /x diameter. Didymium affine, Raunk., Myx. Dan., p. 109 (in English), Tab. 5, figs. 8, 4. On germinating seeds in laboratory. Copenhagen. An unusual Didymium certainly in the wall being without lime. Possibly abnormal. Didymium Fuckelianum. Rost. Sporangia hemispherical, umbilicate below, stipitate, stem whitish with gilvous or brownish shades, strongly plicate longi- tudinally, lime on the wall in the form of stellate crystals, after removal of the lime the wall is covered with unsymnietrical Irown or violet s]-)ots separated ly colourless veins; stem entering the sporangium as a distinct clavate or ap)planatc hroivnish columella. Didi/iuidiH. 223 not at all adherent to lower fart of the sjyoraiif/ial wall, mrface covered with jyrotuherances which give origin to the threads of the capillitium, threads violet, with numerous irregular pro- tuberances; spores lilac, delicately toarted, 9 — 11 fx diameter. Dichjmimn Fuckeliamim, Kost., Mon., p. 161, f. 134; Cooke, Myx. Brit., f.l34; Sacc, Syll., No. 1307. On twigs and pine leaves. Germany. (Rostafinski's Synonyms.) Didymiuin squamulosum, Fckl., Symb. Myc, p. 341, non A. et. Sz. (1569). Didymium Fuckelianum, Rost., Fckl., S. M., 2, Nach., p. 73 (1873). Didymium praecox, De Bary. Sporangia irregularly hemispherical, wall double, the outer white, strongly rugulose, after maturity breaking away in small colourless patches, covered with minute stellate crystals; inner wall greyish lead-colour, rigid, with small irregular granules of Hme, yellowish flesh-colour after removal of the lime; stem generally short, yelloioish fiesh-colotir, with delicate oblong striations, passing into the iJeridiuvi as an irregularly spherical, often applanate yellowish flesh-coloured p)rolongation ; threads of capillitium very delicate, solid, colourless, or some violet with many irregulaily spherical protuberances below the acute angles of the bifurcations, combined constantly by the branches and transverse threads to form a network ; spores subviolet, mem- brane very rigid, with a few scattered short spines, 8 — 9 /x diameter. Didymium praecox, De Bavy, in Rab., Fung. Eur., n. 307 ; Rost., Mon., p. 103 ; Sacc, Syll., 1306. Germany. Didymium squamulosum, Fr. (figs, 40 — 52). Sporangia subglobose, slightly flattened below, and more or less umbilicate, stipitate or sessile, coat of lime at first consist- incc of a continuous white crust of minute cfrauules, which soon 224 A Monograp]( of the Myxogastres. becomes hrokcii wp into large, irregular, persistent patches ; mass of spores brownish-black ; stem ivJdte, equal in length to spo- rangium or much shorter, slightly rugulose ; eolumella ivhite or dirty ochraccous, large, suhglohose ; threads of capillitium pale or colourless, about equal throughout their length, 2 — 3 /x thick, branching at acute angles and connected laterally, forming a net, sometimes flexuous ; spores globose, dingy violet, minutely verrucose, 8 — 11 /x diameter. Diclymium hypnophilum, Mass., Journ. R. Micr. Soc, v., 5. p. 757. Didymium ncglectum. Berk., hiun. Soc. Journ., v., 14, p. 83; Sacc, 1325. a. genuinum. Sporangia stipitate, stem elongated or short, and almost lost in the umbilicus of the sporangium. ^. scssilis. Sporangia sessile, distinct, or sometimes several confluent, when tlie columella becomes elongated. Didymium squamulosum, var. genuinum,, Rost., Mon., p. 159, f. 148; Cooke, Myx. Brit., p. 33, f. 148; Schroeter, p. 122; Raunk., p. 88; Sacc, SylL, 1301. Exsicc.—^y±, Myc. March., 1387, 1388; Cke., Fung. Brit., Ed. II., 613; Rab., Fung. Eur., 813; Rab.-Wint., Fung. Eur., 2969; Roum., Fung. Gall, 1212; Fuckel, Fung. Rhen., 2497; Syd., Myc. March., 396 (as Didymium farinaeeuni) \ and 492 (as Didymi^im herlarum, Fr.); Ellis, N. Amer. Fung., 1216 ; Roum., Fung. Gall., 1684 (as Didymium farinaeeum) ; Klotzsch (Rab.), Herb, Myc, 455 (as Didymium liemisphericum). On rotten wood, leaves, moss, &c. About 1 mm. high, distinguished by the calcareous crust of the sporangium becoming broken up into large irregular patches, separated from each other by the blackish inner layer and the pale columella. Most closely allied to Didymium virgineum. For distinctive characters see under the latter. Britain (Kew, Wothorpe, King's Cliffe, Norwich, Scarboro', Carlisle, Glamis, Appin, N. B.) ; France ; Italy ; Germany ; Switzerland ; Belgium ; U. States ; Cuba ; Ceylon ; Australia ; Tasmania; New Zealand. Didymium. 225 Var. costcdum. Sporangia covered uniformly with minute particles of lime, not hcGoming broken u^ into large, distinct 2>tttchcs ; stem white, variable in length, expanding at the base into a small hypo- tliallus with prominent radiating ridges. Stem sometimes very short, sunk in the umbilicus or entirely absent. Didynvium squaviulosum, /3. leucoims, y. costatum, Rost., Mon., p. 159; Cooke, Myx. Brit., p. 33; Sacc, Syll., no. 1301. Exsicc, — Rab,, Fung. Eur., 367 (as Didymitiiii jJi'uecox, De Bary) ; Fuckel, Fung, llhen., 2690 (as Didymium squamulosum y. costatum) ; Rab.-Wint., Fung. Eur., 2675 (as Didymium Nca- polifanum, Ces.). On leaves, wood, &c. Distinguished from the type by the white crust of the pileus not becoming broken up into patches, and by the markedly costate hypothallus. Somewhat i-esembling Didymium. dqnrs- sum, Fr., which however is readily distinguished by the much larger and more coarsely warted spores. Britain (Epping Forest, Chiselhurst, Kew, Carlisle) ; France ; Germany; Austria. (Rostafinski's Synonyms.) Bcticularia hemispherica, Bull., p. 93 (1791). Diderma sqtiamulosum, A. and S., t. 4, f 5 (1805). Didymium glohosum, v. stipitatiim, Schwarz., Ac. Holm., p. 114 (1815). Zicca stipitata, D. C, Fl. Fr., No. 670 (1815). Tuhulina 2)ediccUaf.a, Poir., Ency., v., p. 373. Cionium fariimceum, Spr., Syst., iv., 528 (1827). Didymium herhantm, Fr., S. M., iii., 120 (1829). Didymium kucojms, Fr., S. M., iii., 121 (1829); Cooke, Hdbk., No. 1127. Didymium costatum, Fr., S. M., iii., 118 (1829). Didymium Ikcoidcs, Duby, Bot. Gall., ii., 864- (1830). Duhjmiitmfilamndosnm, Wallr., Fl. Germ., No. 2187 (1833). Q 226 A MonograpJi of the Alyxogastres. Didymium squ.nimUosnm, Fr., S. M., iii,, 118; Eng. Fl., v., 312; Cke., Hdbk, No. 1122. DifJymium ncghctum, B. and Br., Ceylon Fungi, No. 747. Didymium aastralc, Berk., Hdbk., Flor. Nov. Zealand, p. 191. Var. virgincnm, Mass. Scattered, or in clusters of two or three ; sporangia subglobose, flattened and slightly umbilicate below, external crust of lime pure white, rugulose and crumjdcd but continuous; stem lohite, stout, equal to sporangium or shorter, expanding at the base into a small wrinkled hypothallus ; mass of capillitium and spores blackish; columella siihglohosc or hemisjyJicrical, white or ycllmvish, rugulose; threads of capillitium copious, radiating from the columella to the wall of the sporangium, 3 — 4 jj. thick at the base, a little thinner upwards, often flexuous, sparingly branching at an acute angle, and sometimes connected laterally, with scattered, small, elliptical swellings, pale dingy violet or brownish, sometimes almost colourless ; spores globose, epispore rather thick, brownish- violet, minutely verruculose, 10 — 13 ii diameter. On leaves, wood, &c. Britain (Scarborough, Carlisle, Epping Forest) ; Italy. (Type in Herb., Kew.) Sporangia 1 — 1"5 mm. diameter. The outer coat of lime is very much crumpled, with prominent irregular ridges, but does not become broken up into dctaclicd particles as is usual in the genus Didymium. Didymium microcarpon, Rost. (hgs. 44, 45). S'porangia glohose, at first with an external continuous white crust of lime, which soon becomes broken up into minute glistening particles, resting on the inner dark membrane, more or less umbilicate below, stein slender, slightly attenuated ufumrds, blackish or rufous, longitudinally rugulose, expanding at the base into a small subcircular hypothallus, straight or slightly curved above, generally cd)o^it twice the length of the s^iorangium ; Didipniinn. 227 colniiiclla small, sj^heriml, 2>(d<' ochrcremis, internally traversed by irregular strands giving it a cellular appearance, cavities containing crystals of lime ; capillitium radiating from the columella to the wall of the sporangium, threads about equal, 2 — 3 jj. thick, repeatedly forked and joining laterally, sometimes flexuous and rough in parts with minute granules of lime, varying from colourless, through pale brown to dirty violet; spores globose, very minutely verruculose, brownish-purple, 5 — 7 fx diameter. a. nif/ripes. Stem blackish. /3. rnfi2')cs. Stem rufous or yellowish. Didymium microcarpon, Rest., Mon., p. 157, f. 138, 177 ; Cooke, Myx. Brit., p. 82, f. 133, 177; Sacc, Syll, 1310 ; Eaunk., p. 87. Exsicc. — Ellis, N. Amer. Fung., 1893 (as Didymium micro- mrpmi, Fr.), var. nigri'pcs ; Ellis, N. Amer. Fung., 412 (as Didymium xanthoinLs). On twigs, leaves, living mosses, &c. Plants 2-5 — 3 mm. high, lumps of lime on the sporangium in the form of stellate crystals; umbilicus at base of spor- angium sometimes very slight; columella small but rather prominent; the capillitium varies considerably, sometimes copious, anastomosing, various, inclined to be rough here and there with minute granules of lime, or even nodulose ; at other times scanty, colourless, and threads very thin. So far as I have had an opportunity of ascertaining, the pale-stemmed form is confined to living mosses. Britain (Kew, King's Cliffe, Carlisle, Shere, Forden, Edin- boro', Glamis, N. B.) ; Sweden ; Denmark ; France ; United States; Ceylon. (Rostatinski's Synonyms.) Lyca'pcrdon stiintatuin, Retz,, Vet. de Handl. (1769). Triehia hemispheric a, Trent., p. 228 (1797). Physarum nigripcs, Lk,, Diss., 1, 27 (1809) ; Ditm., t. 42. TricJiia alba, Purt., Midi. Fl., iii., 1113 (1817). Cioniicm xantho]ms, Ditm., t. 43 (1817); Nees, f. 106. Physaruvi 7nicrocarpon, Fr., Gast., p. 23 (1818). 228 A Monograph of the Myxogastres. Didymium lolatutn, j3. stipitatmn, Somm., Fl. Lapp,, 210 (1825). Didijmium nigripes, Fr., S. M., iii., 119 (1829); Eng. FL, v., 313; Cooke, Hdbk., No. 1124. Difhjmiuvi xantJiopus, Fr., S. M., iii., 120 (1829); Berk., Ann. N. H., iii. ; Cooke, Hdbk., No. 1126. Difhjmmm iridis, Fr., S. M., iii., 120 (1829). Didymium micToccpuhaluiii, Chev., Byss., f. 11 (1837). Diilymium melanojms, Wallr,, Fl. Germ., 2184 (1837). Didymium WallrotUi, Eabh., Fl. Cr., 2289 (1844). Didymium porphyroiyics, D. R. and M., Fl. Alg., 409 (1846). Didymium megalosporum, B. and C, Grev., No. 318. Didymium macrospermum, Rost. Sporangia spherical or hemispherical, very much fiattencd vertically, umbilicate below, grey, stipitate, membrane of spor- angium colourless or pale brownish; stem generally longer than peridium, dilated at the base into a distinct discoid hypothallus, attenuated above, strongly plicate longitudinally, snmv-white or yellowish-white; columella versifonn, discoid, &c., threads of capillitium shnple, rarely branching at acute angles, colourless or pale brown ; spores subviolaceous, strongly aculeate, 12 — 13 ju diameter. Didymium macros^ierinum, Rost., Mon., p. 166; Schroeter, p. 122; Sacc, Syll., n. 1304. On trunks. Germany. Gregarious; stem three or more times longer than the sporangium, thinner above, much wrinkled, ridges membran- aceous, pellucid, peridium thin, sometimes dehiscing by be- coming ruptured at the base, and almost all disappearing; at other times torn into shreds which persist. Appears to be allied to Didymium squamulosum, var. costatum, and distin- guished by the flattened sporangium, long stem, and larger spores. (Rostafiuski's Synonyms.) (?) Trichia alata, Trent., 1. c, p. 228 (1797). Didym'inm. 229 (?) Physarum alatum, Fr., 1. c, iii., p. 132 (1829). Diclymium costatum, Fckl., Syni. Myc. I., Nach., p. 339. Didymium discoideum, Rost. Sporangia dueoid, almost sessile, umbilicate, inferior part of wall violet, superior spotted, with irregular pellucid veins, breaking away after maturity in small patches ; stem from its shortness scarcely apparent, hid in the iimhilicus, dilated at the base into a circular hypothallus; columella discoid or hemi- spherical, like the stem yellowish Jlesh-colour ; threads of capil- litium flexuous, rarely forking, bay, tips colourless; spores strongly wartcd, blackish-violet, 11 — 13 ju diameter. Didymium discoideum, Rost., Mon., p. 162; Sacc, Syll., no. 1305. Germany. * * SpiOTCs smooth. Didymium radiatum, B. and C. (figs. G8 — 72). Scattered, sporangia vertically compressed, not umbilicate, but sometimes slightly depressed below, dark grey, with crowded, small, white, innate patches of lime, dehiscing irregularly ; stem longer than height of sporangium, slightly attenuated upwards, white or pale grey, expanding into a smcdl, circular hypothallus, longitudincdly costctte, the ribs passing in a radiate manner into the hyjjothallus ; columella brownish, much com- pressed; capillitium abundant, radiating from the columella to the wall of the sporangium, tlireads thin, brownish-lilac, some- times forked; spores globose, lilac, smooth, 8 — 10 ix diameter. Didymiuiih radiatum, B. and C, Journ. Liim, Soc, vol. x., p. 348; Sacc, Syll., n. 1328. (Type in Herb. Berk., Kew, no. 107G5.) Didymium commutahile, B. and Br., Journ, Linn. Soc, v. xiv., p. 83; Sacc, Syll., n. 1300; Rost., App., p. 21. (Type in Herb. Berk., n. 10766.) Didymium botryoidcs. Berk, in Herb. /. gcniduum. Simple, scattered ; stem usually lunger than sporangium. 230 A Monogi'a2)1i of the My,wgastres. f. fasciculatnm. Three to five sporangia confluent, supported on a common stem. On wood, leaves, moss, &c. Cuba ; N. Zealand ; Ceylon. Remarkable for the circular, sharply limited hypothallus with prominent, regularly radiating ribs, resembling the hy- menium of a small agaric. Resembling superficially some forms oi Didymium sqummdosum, var. costatum. Didymium clavus, Rost. (figs. 53, 54). Sporangia very 'much depressed, slightly convex above, plane below, lime sprinkled in tine crystals on the surface of the dark membrane ; stem short, Uacldsh-hroivn, more or less longitudinally rugulose ; columella absent ; mass of spores blackish ; threads of capillitium 3 — 4 fx thick at base, slightly tapering, sparingly forked at acute angles, and sometimes laterally connected and furnished with small ring-lilce dark-coloured thicl'enings, pale brown to dirty violet ; spores globose, smooth, dingy lilac, 6 — 8 fx diameter. Didymium clavus, Rost., Mon., p. 153; Cooke, Hdbk., p. 3U ; Raunk., p. 87; Schroeter, p. 121 ; Sacc, Syll., 1299. On twigs, leaves, &c. Britain (Wey bridge, Kew, Otton Wood, Leicester ; Epping Forest, Forden, Linlithgow) ; France ; Bel- gium ; Germany ; United States ; Canada ; Egypt ; Ceylon. Sporangia scattered, 1 — 1*5 mm. across wlien circular, some- times irregular and larger. Remarkable for the much flattened, pileus-shaped sporangia. (Rostafinski's Synonyms.) lieticularia hemisphcrica, Bull., t. 446, f 2 (1791). Physarum clavios, A. and S., t. 2, f. 2 (1805). Didymium melanop^ts, j8. clavus, Fr., S. M., iii., 114 (1829); B. and Br., Ann. N. H., No. 110 ; Cooke, Hdbk., No. 1118, /3. Didymium hemiq)hericum, Wallr., Fl. Germ., 2192 (1833). Didymium clavus, Rabh., Fl. Crypt., 2282 (1844). Didymium clavus, Berk., Eng. FL, v., p. 314. Dklym'uim. 231 Didymium neglectum, Mass. (nov. sp.) ; (tig. 207). Sporangia globose or slightly depressed, flattened or some- times slightly umbilicate below, stipitate ; wall single, at first with a continuous, rugose, white crust of lime, which at maturity becomes broken up into minute, persistent particles; stem equal to or longer than height of sporangium, erect, slightly attenuated upwards, hlaclx, longitudinally wrinkled, expanding at the base into a small hypotballus ; columella circular in outline, much dcjircssed, placcntifoi'm, with a distinct margin, covering the base of the sporangium, rugulose, imle hrownish- lilac, capillitium dense, threads radiating from the columella, lilac-brown, becoming colourless towards the tips, bifurcating, now and then anasmosing laterally; spores globose, smooth, 7 — 9 IX diameter. On wood. Fairmount Park, Philadelphia, U. S. Solitary or in scattered clusters of 2 — 6 specimens, about 1 mm. high ; superficially resembling Lidymiuvi farinaccu.m, but quite distinct in the smooth spores, which only show vague indications of roughness when magnified 1200 diameters. The columella is very characteristic, being broad and flattened with a distinct margin ; at maturity the sporangium falls away except a minute frill round the lower edge of the columella, which, with the stem, in this condition resembles a little table. The original specimens were found mixed with specimens of I'ihiiadoclic ohloiuja, communicated by Mr. Harold Wingate, of Philadelphia. Didymium Barteri, Mass. (n. sp.). Gregarious; sporangia sphcrico-depressed, slightly umbilicate below, stipitate, cretaceous layer pure ivhitc, rugulose, for a long time persistent, then breaking away in small granules; stem elongated, slender, attenuated uincards, jj76?-(! white, strongly wrinkled longitudinally; columella small, globose, u'At^^c; mass of spores blackish-purple ; capillitium copious, threads about 2 ju thick, equal, sometimes floxuous, forked at an acute angle 232 Monograph of the Mijocotjastres. and combined laterally to form a flaccid, irregular net ; spores globose, smooth, pale dingy lilac, 10 — 11 }x diameter. (Type in Herb., Kew.) On moss. W. Africa (Niger Expedition, Barter). A very beautiful and distinct species, known at once by the snow-white sporangium, stem, and columella, and the smooth spores. Plant 3 mm. high; stem 2-5 mm. high, sporangium •5 mm. high, a little more in diameter. B. Sporangium sessile. (In B. sjntmarioidcs and D. 2>hi/sarioides, a very short stem is sometimes present.) Didymium Alexandrowiczii, De Bary and Rost. Sporangia sessile, defm^med, not truncate, several often collected in little clusters, rugidose, granulose ; columella ahsent ; threads brownish, colourless at both ends, simple or bifurcating and laterally connected; sjjorcs smooth, obscure violet, 10 — 12 jut diameter. Didymiu'ia Chomlroderma, De Bary and Rost., in Alex. Stroz,, p. 89. Chondrioderma Alexcmdrmviezii, Rost., Mon., p. 1G9, f 17G ; Sacc, Syll, 1253; Cooke, Myx. Brit., f. 176. On wood. Britain (Kew) ; Poland. Didymium spumarioides, Fr. Sporangia substipitate or sessile, scattered or densely crowded on a well-developed, white or imle fiesh-coloured hypothallus con- taining much lime; wall of sporangium white or grey, crust of lime ^ndvcrulent ; columella variable, smooth or irregularly rugulose, white or pale flesh-colour, sometimes almost obsolete ; threads of capillitium about equal, 1"5 — 2 ju, thick, repeatedly bifurcating and connected laterally to form a net, colourless or dingy lilac, spores globose, minutely warted, 9 — 12 ju diameter. Didymium spumarioides, Fr., Symb. Gast., 20. Chondrioderma sjmmarioidcs, Rost., Mon., p. 173, f. 142 — 145, Diih/mmw. 233 151; Cooke, Myx., p. 38, f. 142—145, 151; Schroeter, p. 123 ; Sacc, Syll., n. 12G9. Chondriodcrma stroviateum, Rest,, Mon. Append., p. 18, f. 151. Didcrma farinaccnni, Peck, 26tli Report State Mus., N. Y., p. 74. Exsic.c. — Fckl., F. Rlien., 2495 (as Carccrina spuntarioidrs, Fr.) ; Rab., F. Eur., 432 (as Carccrina sjnwiarioidcs, Fr.) ; Cke., F. Brit., Ed. II., 825 ; Roum., Fung. Gall., 1682. On living or dead leaves, moss, &c. Britain (Higligate, Shore, Scarboro') ; France ; Germany ; Sweden ; Belgium ; Italy ; United States ; Australia. The present species was placed by Rostafiuski in the genus Chondrioderma, from which it differs, according to our inter- pretation of the genus, in the pulverulent, instead of porcelain- like coat of lime. Very variable in colour and size of the columella, but well marked by the highly-developed hypo- thallus, which sometimes becomes so thick as to suggest the idea of a stroma. Sporangia from '5 — 1"5 mm. diam., circular or irregularly angular from mutual pressure, sometimes the stroma, when very thick, forms a short stem-like projection ; the colour of the stroma and columella varies from chalk-white through pale tints to pale flesh-colour. (Rostafinski's Synonyms.) Sjnomaria physarioides, Pers. Syn., 163 (18U1). Fhysarum didcrmoides, Fries, Herb. Spumaria alha, Schum. Saell., No. 1414 (1803); Fl. Dan., t. 1798, f. 2. Didymium spiumarioidcs, Fr., Symb. Gast., 20 (1818). Fhysarum stromatium, Link, Hdbk., iii., 409 (1833). Carccrina sp'amarioides, Fr., S. V. S., 451 (1849). Didcrma sjmmarioidcs, Fr., S. M., iii., p. 104; Eng. FL, vol. v., p. 311; Cke., Hdbk., No. 1109. Didymium physarioides, Rest. Sporangia cylindric-ohlonff, often crowded and ilefornied, seated on a well-developed hypothallus, sessile or with a short 234 A Mo7W(ji'aph of the Mij-xogastres. stem, wall dark grey, at first with granules of lime ; coluinella irregular, more or less confluent and common to all the sporangia ; threads of capillitium 2 — 3 fi thick at base, with but few bifurcations, with mimerous fusiform or discoid dingy 'pmylc thichcnings, spores globose, brownish purple, vmrtcd, 12 — 14 fx diameter. Didymium ]ihysarioidcs, Rost., Mon., p. 158, fig. 147; Cooke, Myx. Brit., p. 33, f. 147; Sacc, Syll., no. 1811. On moss and wood. Britain (King's Cliffe, Carlisle); Ger- many ; Sweden ; United States ; Cape of Good Hope. Often superficially resembling Fhysarum didermoides. Didymium serpula, Fr. (figs. 55 — 57). Sxmrangia sessile on a hroad base, much addressed, circular in outline, or variously elongated and flexuous, and sometimes the branches combine to form an irregular network, wall dark grey, sparingly sprinkled with minute luhite jJarticles of lime ; columella absent ; mass of spores blackish ; threads of capillitium branched and connected laterally, forming an irregular network, dirty brown to colourless, plentifully furnished with dark-coloured, riug-like, or irregularly-shaped thickenings; spores globose, very mimitely verricculose, 7 — 9 fj. diameter. Didymium serpula, Rost., Mon. Append. ; Cooke, Hdbk., 30, figs. 166, 180 ; Schroeter, p. 121 ; Sacc. Syll, no. 1297. Didymium complanatum, Rost., Mon., p. 151, figs. 166, 180. On leaves, &c. Sporangia when circular -5 — 1 mm. diameter ; when elongated often several mm. in length. Sometimes the lime on the sporangium is more abundant, forming a white crust ; but the species cannot be mistaken if attention is paid to the much flattened sporangia, the well-developed capitulum having the threads furnished with numerous, variously-shaped, dark browu thickenings, and the very minutely verruculose spores. Britain (Kew, Batheaston, Scarboro', Carlisle); all Europe; United States. Didymmm. 235 (Rostafiiiski's Synonyms.) Lycopcrdon comi^hniatum, Batscli, t. 170 (178C). Didymmm scrpida, Fr., S. M., iii., 126 (182!)); Eng. Fl., v., 314; B. and Br., Ann. N. H., No. 1035; Ckc, Hdbk., n. 1134. Physarurii conjlnem, Fckl., Symb. Myc, p. 342 (1809), non Pers. Didymium confluens, Rost, Simrangia sessile, sometimes scattered and then hemispherical, at others densely crowded on a well-developed hypothallus, hemispherical reniform, or irregularly angular from mutual pressure; columella distinct, yclloioish, or with a ^nWj tinge; threads of capillitium very slender, forlced at acute angles, ixde hrownish-lilac' ; spores globose, rather coarsely wartcd, dingy purple-brown, 10 — 15 // diameter. Didymium conjliiens, Rost., Mon., p. 164. Didymium erustaceum, Sacc, SylL, no. 1303. On leaves, wood, &c. Britain (Castle Howard, Yorks) ; France; Italy; Russia; Finland. (Rostafinski's Synonyms.) Didymium eomplanatum, Schrad., Nov, Plun. Gen., p. 24, t. 5, f. 5 (1797). Physarum confluens, Pers. Syn., p. 169, et v., a truncigerum (1801). Physarum confluens, /3 muscigcnum, A. et Sz., 1. c, p. 9 (1805). Didymium erustaceum, F., 1. c, iii., p. 124; Excl. Syn. (1829). Cionium complanatum, Lk. ap. Wallr., 1. c, no. 2176 (1833). Var. ohducens, Karst., Not. Sallsk., pro. faun, et flor. Fenn., ix., p. 356 (1880). Sporangia irregular (plasmodiocarp), flattenetl, often umbili- cate, concave below, solitary or gregarious, sometimes densely crowded ; stem short or almost obsolete, irregular, broadening out below into the common hypothallus. Remainder as in type. 236 A Monograph of the MyxogaMres. Didymium eflFusum, Rost. Sporangia densely aggregated or scattered, sr.ssi/e on a hroad hasc, circular or irregular in outline, dcjyrcsscd, wall grey, pruinose with numerous minute crystals aggregated in masses; columella hemis2oherical, white or pallid, rugulose ; threads of capillitium very slender, colourless or pale inXiguxoxxs,, furnished with scat- tered coloured protuherances, combined to form an irregular dense net; spores globose, brownish-purple, minutely warted, 10 — 12 ju, diameter. Didymium effusum, Rost., Mon., p. 163; Sacc, Syll, no. 1302. On leaves, wood, &c. Britain (Cornwall) ; Finland ; Sweden ; France ; Germany ; United States ; Ceylon. About '5 mm. diameter, characterized by the white columella, the capillitium threads furnished with numerous well-defined projections in the form of thickened plates or pyriform swellings, and the dense meal of crystals on the wall. (Rostafinski's Synonyms.) Diderma effusum, Link, Obs. I., p. 42 (1809). Didymi'um effusum, Fr., 1. c, iii., p. 124 (1829). Pliysarum conjlucns, Fr., Excl. Syu., 1. c, iii., p. 146 (1829). Sub-Gen. Hemididymium . Capillitium usually dense, threads branching and anastomosing to form an irregular network throughout the capillitium, angles usually flattened and more or less triangular. The species included in the present sub-genus are distin- guished from those belonging to the sub-genus Didymium by the absence of the simple or bifurcating threads at the basal portion of the capillitium. A. Siwrangium stipitate. * Sjjores smooth. Didymium longipes, Mass. (n. sp.) (fig. 226). Sporangia small, globose, snoiv-v:hitc, frosted with a few scattered granules or crystals of lime ; stem very long and Didyiuiitm. 237 slender, erect, snoiv-whitc, very slightly attenuated upwards, almost smooth, expanding at the base into a small, circular, white hyiwthallus ; columella absent ; capillitium well-developed, threads very thin, colourless, hranching and anastomosing irregu- larly to form a network, nodes usually triangular; spores globose, dingy hlac, smooth, 8 — 10 /u, diameter. (Type in Herb. Berk., Kew, with Tilmadochc nutans.) On bark and wood. Britain (Yorks) ; Lower Carolina. A very distinct species, characterized by its snow-white colour throughout, 2'5 — 3 mm, high, sporangium very small. Spores perfectly smooth under a yV objective. Didymium fulvellum, Mass. (n. sp.) (fig. 232), Sporangia spherical, very little or not at all umbilicate, stipitate, outer cretaceous coat pure white, co'mposcd of minute granules which fall away in the form of ijowder, inner wall with- out lime, rather tough, bright fulvous; stem slender, elongated, straight or slightly curved above, attenuated ujnvards, same colour as inner wall of sjiorangium,, much wrinkled longitudin- ally, expanding at the base into a similarly coloured circular liypothalhts ; columella broicn, not very prominent ; mass of spores blackish with a purple tinge ; capillitium copious, threads repeatedly forked, 2 — 3 /u., thick, rather broader at the base of each bifurcation, combined laterally to form an irregular net- work, colourless; spores globose, smooth, pale dirty lilac, 8 — 10 \x diameter. On fallen oak leaves ; scattered or gregarious. United States. (Type in Herb,, Kew.) Sporangium 1 mm. diameter, stem about 2 mm. high, Didymium australis, Mass, (fig, 208), Sporangium globose or slightly flattened above and below, slightly umbilicate, at first with a thick, white la^^er of crystals which soon brealcs up into large, 2'>crsistent, irorgnlar patches; stem thill, hrig/tt Irovm, slightly attenuated upwards, often bond- 238 A Afonograph of the Myxoga,9tres. ing under the weight of the sporangium ; columella alscnt ; mass of spores blackish-purple ; capillitium copious, threads colourless, 3 — 4 fi thick, branching at wide angles and anasto- mosing irregularly ; spores globose, smooth, dingy lilac, 9 — 10 ju diameter. Didyniium australis, IVIass., Grev., vol. xvii., p. 7. (Type in Herb., Kew.) Gregarious, on an old decayed species of Auricularia. Brisbane, Australia. Plant about 2 mm. high, stem equal or a little longer than sporangium. Resembling Didymium sqtiamnlosum, in the wall of the sporangium breaking up into large detached patches, but readily distinguished by the absence of a columella, and the tliin, coloured stem. Didymium proximum, B. and C. (figs. 227 — 229). Sporangium globose, white or pallid, sprinkled with lime, mostly in the form of stellate ci'ystals ; stem more or less longi- tudinally wrinkled, suhequal, rufous, erect, expanding at the base into a small hypothalhis; columella suhclavate, ivhitish, reaching one-third the height of the sporangium; capillitium dense, threads colourless, sometimes rough with projecting granules ; 3 — 4 mm. thick at the base, bifurcating and anasto- mosing laterally to form an irregular reticulation, often dilated and flattened at the axils; spores globose, pale dirty lilac, smooth, 9 — 12 fx diameter. Didymium proximum, B. and C, Grev., vol. ii., p. 52 (1873) ; Rest, Mon. App., p. 23; Sacc, Syll., no. 1308. Didymium ptisillum, B. and C, Grev., ii., p. 53. On dead pine leaves and on herbaceous stems. Carolina. (Types in Herb. Berk., nos. 10760 and 10764.) Scattered, 2 — 2'5 mm. high, columella large, usually more or less clavate, sometimes irregularly fusiform. Owing to a slip of the pen, Berkeley says, in describing D. pusiUum, " columella nulla," yet the large columella is shown in Berkeley's sketch, and the type specimen is identical with that of D. p)ro:nmum. Didyminm. 230 Spores smooth, or with sometimes the shghtest suggestion of gramilatiou under a -pV objective. Didymium pezizoideum, ]\Iass. Sporangia stipitate, discoid, ijczizacfonn, greyish-white, Avail single, hccoming fnrfuraceous ; stem subulate, curved above, brownish, smooth, expanding at the base into a small, shining, closely adnate hypothallus ; threads of the capillitium combined to form a net, spores smooth, violet, 8 — 9 /x diameter. Trichampliora ijczizoidea, Jungh., Fl. Cr., Java, p. 12, t. 2, f. 9. Chondrioderma2)ezizoidcu'm,'Ro&t.,'Mon.,Tp. 424, f. 122; Cooke, Myx. Brit., fig. 122; Sacc, Syll., 1256. On rotten trunks. Java. Didymium platypus, Hanzl. Greyish-white, primrose, sporangia scattered, convex above, deeply umbilicate below ; stem cylindrical, dilated in a discoid manner at the aj^cx ; spores globose, smooth, blackish, 8 n. diam., capillitium scanty, formed of black tubes connected in a reticulate manner, columella absent. Hanzl. Ein neue Myxog. typus in Just. Bot. Jaliresbericht, 1872, p. 155; Sacc, Syll., 1326. Agaricus platypus, Bischoff (sec. Hanzl.). On putrid plant stems. Hungary. * * Spores icartcd. Didymium echinospora, Mass. (n. sp.). Sporangia globose, usually with a shallow umbilicus below, wall membranaceous, almost colourless, with a dense coat of white amorphous particles of lime, upper portion breaking away in flakes during dehiscence, and leaving a permanent cu2^-like hasal portion with torn edges; stem elongated, slender, erect, very slightly attenuated upwards, white, rugulose below ; colu- mella absent; capillitium very dense, threads lilac-brown, about 2 /x thick, hifurccding, and towards the apex freely anastomosing 240 A Monograph of the Myxogastres. to form a dense net, with the angles usually triangular ; spores dingy purple, globose, loartcd, 12 — 14 ^ diameter. (Type in Herb. Berk., Kew., with Tibnadodie nutans.) On bark. Ceylon. Scattered, 1'5 — 2 mm. high. Bearing a superficial resem- blance to Tilmadoche nutans, but distinct in the capillitium, which contains no trace of lime, and in tlie spores. Didymium zeylanicum, B. and Br. Sporangium hemispherico-depressed, with a shallow umbilicus below, wall thin, colourless, frosted with white granules of lime ; stem elongated, attenuated v^nvards, or almost obsolete, brovn or yellow, longitudinally wrinkled, filled with small grains of lime, expanding below into a spreading hypothallus ; capillitium forming a dense irregular net, threads colourless, very thin ; spores dark purple, globose, very minutely verruculosc, 7 — 9 ju diameter. Didymium zeylanicum, B. and Br., in Journ, Linn. Soc, vol. xiv., p. 84. Chondrioderma zeylanicum, Bost., Mon. App., p. 15 ; Sacc, SylL, no. 1254. Chortdrioderma. exiguwm, Racib., Hedw., 1889, p. 118. (Type in Herb. Berk., no. 10770.) On decayed Avood. Ceylon ; Poland. The species described as Clwndrioderma cxiguum cannot be separated from the present species; the points of difference given by Raciborski only apply to certain individuals which, from an examination of the type, is a variable species. Didymmm obrusseum, B. and C. Sporangium white or pale yellow, almost globose, sometimes slightly umbilicate below, wall very thin, ivith little or no lime, often iridescent when empty ; stem attenuated ujnvards, fattened and distorted helow, weak and droojnng, hrigJif yelloio, expanding at the base into a very thin, irregular hypothallus of the same colour ; columella absent ; capillitium rather scanty, threads Didyminm. 241 thin, colourless, repeatedly forked and joined laterally to form a loose, irregular net ; spores globose, dingy lilac, mry minutely verruculose, 8 — 10 [x diameter. Didyminm ohrusscum, B. and C, Grev. (1873), vol. ii., p. 53. Physarum ohrusscum. (B. and C), Sacc, Syll., 1195 (in part). (Type in Herb., Kew.) Exsicc. — Fung. Cubenses Wrightiani, 532. On twigs, leaves, &c. Cuba ; United States. Gregarious, stem flaccid, irregular and flattened below, very thin above, about 2 mm. long, hypothallus very thin and shining. Didymium pertusum, Berk. Scattered. Sporangium globose or depressed, generally more or less umbilicate, u^hitc, frosted with minute crystals of lime ; stem elongated, sliglitly attenuated upun7ds, rufous; columelki entirely absent ; threads of capillitium very thin, colourless, vari- ously branched and combined to form a net ; spores globose, dirty lilac, verruetolose, 10 — 13 /x diameter. Didymium pert'USiLm,^exk., Eng. Fl., vol. v., p. 313; Cooke, Myx. Brit., p. 35; Sacc, Syll., 1317. (Type in Herb. Berk.) On dead herbaceous stems. Appin, N. B. Scattered, varying from 2'5 — 3 mm. high. There is no trace of a columella, although after dehiscence the basal persistent portion of the sporangial wall falls down, and this under a pocket-lens might be mistaken for a columella. The threads of the capillitium anastomose freely and form a very irregular, scanty network. The stem is more or less longitudinally rugulose, and filled with amorphous particles of lime. Didymium eximium, Peck. Sporangia subglobose, slightly or not at all umbihcate below, grey, frosted with stellate crystals of lime ; stem elongated, erect or slightly curved above, almost equal, slender, i^alc yelloioish- hrourii, darker towards the base, which expands into a minute hypothallus, slightly rugulose, filled with amorphous particles 242 A Monograph of the Myxogastves. of lime, as is also the orbicular, depressed, jJCf'l^ columella; capil- litium colourless, threads very slender, combined into a dense ii'vegidar net ; spores globose, brownish-lilac, miimtely loartcd, 8 — 11 \x. diameter. Didymium eximiu)ii, Peck, in 81st Rep. N. York State Mas., p. 41 ; Sacc, Syll., n. 1314. Uxsicc.—Ellis and Everhart, N. Amer. Fung., Ser. II., 2089. (Authentic specimen from author in Hb., Kew.) On dead leaves, hei'baceous stems, &c. U. States. Scattered or gregarious, from 1 — 1"5 mm. high, stem twice or more the length of the sporangium. Threads of capillitium often more or less dilated at the angles, and rarely with a sliglitly thickened interstitial portion, which does not however contain lime. Didymium flavicomum, Mass. (figs. 76 — 78). Sporangium depresso-globose, minutely umbilicate below, pale yellow, at first frosted with a few scattered crystals of lime ; stem elongated, slender, erect or slightly curved near the apex, subequal or attenuated upwards, yellowish-rufous, often darker at the base, which expands into a minute hypothallus, longi- tudinally wrinkled and containing amorphous granules of lime ; columella absent; capillitium pale yellow, threads variously combined to form a rather dense net, dilated at the nodes, which are without a trace of lime ; spores almost colourless, very minutely verruculose, 9 — 10 [i diameter. Physarum fiaviconuLm, Berk., Hook. Journ. Bot., vol. iv., p. m-, Sacc, Syll, n. 1193 (in part). Physarum Bcrheleyi, Host., Mon., p. 105. (Type in Herb. Berk., n. 10,782.) On decorticated wood. Australia (Swan River), Gregarious, 2-5 — 3 mm. high, stem slender, three to four times as long as the sporangium. The present very distinct species has been hopelessly confused by Rostafinski and by Berlese in Saccardo's Sylloge, and the mistakes committed by both can be traced to the modern pernicious system of paying attention to books rather than specimens, always a mistake, and Didymmm. 243 more especially so when dealing with diagnoses of microscopic species described half a century ago. The following synonyms by Rostatiuski, although absolutely worthless from a scientific standpoint, may be historically interesting. (Rostafinski's Synonyms.) Physarum Jiavicomum, B., Hook. Journ., p. QQ, n, 63 (1845), Stylonitcs fulviceps, Fr., Fung. Natal, p. 33 (1848). Physarum mjjriceps, B. et Rav., Fung. Car. Exs. iii., n. 76 (1855). Ophiotheca ? Roussel. Antil. (1872). Physarum cuprijjes, B. and Rav., Grev., p. 65, n. 355 (1873). Physantm rosetcm, B. et Br., cfr. Grev., p. 65 (1873). Didymium elegantissimum, Mass. (n. sp.). Sporangia scattered, stipitate, hemispherico-depressed, with a broad, shallow umbilicus below, smw- white, with a dense coat- ing of stellate crystals; stem elongated, equal, rather stout, longitudinally rugulose, bright orange-hroivn, passing into a small hypothallus of the same colour; capillitium absent; threads 3 — 4 /^ thick at the base, forking at acute angles, here and there anastomosing to form irregular portions of network, most of the main branches with numerous small nodulose siuellings at regular intervals, giving a moniliform appearance, pale and lilac-brown below; spores globose, 8 — 10 /x diameter, mwrni^gZy vMrtccl, pale lilac. Physarum striatum, Fr., Syd. Myc. March., 490 (Kew copy). On twigs and leaves. Britain (Scarboro') ; Germany. About 1-5 mm. high. Stem two or three times as long as sporangium. Bears a superficial resemblance to Didymium p>crtusum, Berk., from which it is readily distinguished by the denser capillitium aud the moniliform thickening of the threads. B. Spoi'angimii sessile. Didymium leptotrichum, Mass. Sporangia venulose or irregular, depressed, wall single. 244 A MoJior/rapk of the Myocogastres. covered with minute sqiiaraules of lime ; columella absent ; threads of the capillitium very slender (08, 3 ju diam.), forming a flaccid, very dense net; spores blackish-brown, verruculose, 12, 5—13, 2 ix diameter. Chondriodervia leptotrichum, Racib., Myx. Crac, p. 7, f 4 ; Sacc, Syll., 1267. Poland. Eemoved from Chondrioderma on account of the furfuraceous and not porcelain-like wall of the sporangium. Didymium Listeri, Mass. (figs. 97 — 101). Plasmodium jndvinato-comprcssed, in large, irregular, scattered 2')atdies ; outer wall white, with a thick layer of stellate crystals of lime, rather fragile, breaking away in irregular patches ; inner wall very thin ; columella absent ; cainllitiwn dense, threads parallel, without free ends, often forming acute-angled dichoto- mies and connected by transverse bars, dark brown, usually branched near the paler tips that are attached to the wall; spores globose, dingy lilac, minutely warted, 8 — 10 /x diameter. " Didymium dulium, Rost.," Lister in Herb., Kew. On dead leaves. Britain (Lyme Regis). The present species superficially resembles D. didnum, but is quite distinct in the copiously anastomosing threals of the capillitium which are entirely destitute of free, curved, spine- like branches, Plasmodia 3 — 7 mm. across. Didymium Tussulaginis, Mass. Sjiorangia sessile on a hroad hase, hemispherical or irregular and plasmodiocarp, wall thin, pinkish-grey, more or less iri- descent, with minute, scattered crystals of lime, dehiscing irregu- larly ; capillitium well-developed, springing from the base of the sporangium, threads repeatedly branching in a dichotomous manner, rarely anastomosing, with scattered, small sivellings usually con- taining one large or two. to four small crystals of lime; spores Uack-lroion in the mass, pale brown with a lilac tinge by transmitted light, globose, very minutely verruculose or quite smooth, 12 — 14 /x diameter. Didymiam. 245 Didymium Coolri, Raiink., Myx. Dan., p. 109, t. 5, f. o. Physarum tussilaginls, B. and Br., Ann. Nat. Hist., n. 1597; Cke., Myx. Brit., p. 16. Chondriodcrvia Coolxi, Rost., Mon. App., p. 17. Exsicc. — Cke., Fung. Brit., Ser. I., 520 (as Badhamia ca]}su- llfcr); Cke., Fung. Brit., Ser. II., 206; Tliura., Myc. Univ., 1100; Vize. Micro-fungi Brit., 1 and 101. (Type in Herb., Kew.) On the under surface of living leaves of colts-foot, Tnssilago farfara. Britain (Cheshire) ; Denmark. A very remarkable species, not agreeing in important points with any established genus, scattered or in groups of 2 — 4, 15 — 3 mm. across. Apparently quite normal in its develop- ment, and not accompanied by any other species, it occurred in abundance for several years in succession in Cheshire, and has also been found in Denmark. In a note attached to the specimens in the Kew Herbarium, Mr. A. Lister says, "The whole appearance conveys the impression that it may not be a stable form, and the habit is so peculiar." Of course it is a matter of opinion as to whether the appearance described above can afford any clue as to stability, but certainly many species occur on living plants. The presence of lime in the capillitium prevents the present species from being a typical Didymium or Chondrioderma ; the lime is too scanty and the capillitium too slender for a Physarum. Didymium anomalum, Mass. Plasmodium venulose, iircuate or variously conturtcd, very convex, surface minutely granulose ; columella very strongly dc- velo2)ed, with a broad base resting on the substratum, arcuate, folloiving the shape of the plasriiodioearp ; threads of capillitium very delicate, thin, colourless, ^ixccvX, forming a dense net ; spores oninutcly verrueulose, yellow, 11 — 13 ju diameter. Chondrioderma anonialwii, Rost., Mon., p. 169 ; Sacc, Syll., 1263. Russia. 24G A Monn(fraj)h of the Miixogastres. Tlic granulosc wall of the sporangimii separates the present species from the genus Chmidriodcnnc. Didymium dubium, Host. (fig. 102). Plasmodium pulvinate, arcuate, in crowded clusters, the outer Avail white, with stellate crystals of lime, hreaking away from inner wall in irregular pieces; threads of capillitium rigid, rarely forking, forming a loose net, fvrniahed with curved sinnes, spores obscure violet, minutely vcrrundosc, 10 — 11 ix diameter. Didyriiium. duhium, Rost., Mon., p. 153; Sacc, Syll., n. 1298. Britain (Lyndhurst) ; Bohemia. Sub-Gen. Atricha. Capillitium obsolete ; columella absent. The two species constituting the present subgenus are un- doubtedly degenerate species of Didymium, characterized by the very rudimentary condition, or entire absence of the capillitium. Didymium sinapinum, Cooke (figs. 167, 168). Clavate, subcylindrical, or more or less irregular, wall brittle with lime, at first compact, then hreaking ujy into bright yellow, easily detached, 2)articlcs, becoming irregularly ruptured at the apex, basal attenuated stem-lilcc 2^ortiun darker, often vAth a ■purjile tinge ; hypothallus spreading, thin ; mass of spores blackish -purple ; spores globose, dingy lilac, minutely warted, 7 — 9 [X diameter. Didymium sincqnnum, Cooke, Brit. Myx., p. 33, fig. 245 ; Sacc, Syll., n. 1319. (Type in Herb., Kew.) On leaves. Wales (Forden). Growing in isolated patches, gregarious or crowded, springing from a common hypothallus, sporangia 1 — 1 '5 mm. high, usually moro or less clavate, now and then irregularly lobed and form- ing a small aethalium consisting of two or three sporangia, bright sulphur-yellow ; in rare instances a few very thin, bifur- Didymram. 247 eating thrccads are present, but iu most sporangia there is no trace of a capillitiuin. Didymium flavidum, Peck. Sporangia glohose, or hroadly obovate, sessile on a broad or narrow base, wall brittle with lime, rugulose, sid'phur-yellow, sometimes with a tinge of orange, becoming irregularly ruptured at the apex, or the apical portion breaking away in an irregu- larly circumscissile manner, basal half persistent ; mass of spores blackish- violet, spores globose, dingy lilac, very mimitely ver- ruculose, 10 — 12 // diameter. Didymium flavidum. Peck, 28th Report State Mus., N. Y., p. 54. Physarum flavidum, Peck, Berl., Sacc, Syll., n. 1215. (Authentic specimen from Peck, in Herb., Kew.) On wood. United States. Scattered or gregarious, about '5 mm. diam., springing from a very thin hypothallus ; capillitium consisting of a few scat- tered, very thin, bifurcating threads, or most frequently entirely absent. Owing to iiivpcrfed diagnoses and absence of type sijccimcns, the folloiving cannot he arranged in their respective sections. Didymium tenerrimum, B. and. C. Sporangium globose, white, venulose, umbilicate below ; hypo- thallus obsolete; stem elongated, thickened below, pallid, hyaline; threads of capillitium colourless ; spores black. Didymium tenerrimuvi, B. and C, Linn. Soc. Journ., vol. x., p. 348. Didymium ohrusseum, Sacc, Syll., n. 1195 (in part). On leaves of grass, nearly allied to D. ohrusseum, B. and C. Cuba. The present species is given as a synonym of D. ohnisseum , B. and C, in Saccardo's Sylloge, but as the authors considered the two species to be distinct, and as no specimen exists in Berkeley's Herbarium, I have considered it advisable not to 248 A Monograph of the Afj/avgastres. combine the two, as the present species appears to differ in the absence of a hypothallus. Didymium Weinmanni, Fr. Sporangia turbinate, very fragile, covered with grey meal, stem short, with a yellow tinge, columella absent ; spores blackish. Didymium Weinmanni, Fr., S. M., iii., p. 121 ; Sacc, Syll., n. 1331. On living herbaceous stems. Russia. Minute, closely gregarious, and often arranged in lines ; stem equal, expanded at the base ; sporangia membranaceous, oblong or pyriform, covered with grey meal. (Fr.) Didymium humile, Hanzl. Sporangia applanate, grey, pruinose, apex slightly, below deeply umbilicate ; stem cylindrical, brown, short; capillitium of simple, tortuous, smooth, brown threads ; spores brown, 6 — 7 /a diameter. Hanzl. Ein neue Myx. types, in Just. Bot. Jahresb., 1872, p. 155 ; Sacc, Syll., 1324. Hungary, Didymium fulvipes, Fr. Stem compressed, sulcate, foxy-vermilion ; sporangium globose, delicately villose, grey. Didymimn ftdvi2KS, Fr., Stirp. Femsj., p. 83 ; Fr., Gast., p. 24 ; Sacc, Syll, 1323. Sweden. Stem and hypothallus resembling those of Hemiarcyria ruhi- formis, very unequal, compressed, subconfluent, longitudinally sulcato-rugose, 2 mm. and more in length ; sporangium globose, or slightly depressed, obtuse, often confluent, with a very delicate grey villose covering ; columella absent ; threads brown. (Fr.^) The villose covering of the sporangium alluded to by Fries would in all probability consist of a very subtle coating of a pulverulent nature. Didymium. 249 Didymium versipelle, Fr. Sporangia lenticular, umbilicate below, at first white, pruinose, becoming polished, chestnut, stem conical, rugose, pallid fulvous ; columella brown ; spores smoky-black. Didymium vcrsixidle, Fr., Syst. Myc, iii., p. 117; Sacc, 1830. Sweden. Stem about 2 mm. long, conical, wrinkled, sporangium de- pressed, umbilicate, wall thin, whitish-pruinose, then naked and shining, chestnut, dehiscing by a longitudinal fissure like a bivalve ; columella large, globose, brown. (Fr.) Didymium Fairmani, Sacc. Sporangia scattered, sessile, with stellate crystals; columella subglobose, brownish ; threads of capillitium hyaline, forming a loose reticulation ; spores smooth, 8 — 10 /x. Didymium Fairmani, Sacc, Journ. Myc, vol. v., p. 180 ; I.e., vol. v., p. 78. On leaves of Smilacina hifolia. Syndonville, Orleans County, N. Y. The diagnosis is altogether inadequate, and it is much to be regretted that the seventeenth century style of description should be resuscitated at the present day. Professor Saccardo's description is thoroughly bad, inasmuch as it does not describe anything. Didymium erythrinum, Berk. Stem equal, vermilion, as is the globose sporangium, spores black, flocci Avhite. Berk., Grev., vol. ii., p. 52 ; Sacc, Syll., 1322. The pulverulent matter with which the peridium is clothed soon rubs off. As there is no specimen of the above species in Berkeley's Herbarium, I am unable to add to the above meagre description. Didymium angulatum, Peck. Sporangia delicate, subglobose, whitish, externally mealy with numerous granules and crystals of lime; stem short. 250 A Mouor/rajy/i of the Myxogastres. whitish ; columella subglobose, white or pale yellow ; capillitium sparse, delicate, whitish or slightly coloured ; spores irregular, angular, blackish in the mass, -OOOSo' — -0005' long. Didymium angulatum, Peck, 31st Report, State Mus., p. 41 ; Sacc, Syll., 1312. Fallen leaves. Adirondack Mountains. Aug. I have no knowledge of the present species except from the above description from Peck. Didymium Paraguayense, Speg. Sporangia at first globose or elHptico-elongated, '5 — 1 x "5 mm., sessile, rarely shortly substipitate, deep violet, siDringing from a brown, mucedinous, very thin hypothallus, then hemispherical or reniform, 1 — 2 mm. long by -8 — 1 mm. high, sessile, sub- repens, wall white, thick, cracked or breaking up in flakes; mass of spores ftdvous or ;pale huff; threads of capillitium very thin, hyaline, not evanescent; spores globose, inrfcctly smooth, pale olive brown, or tobacco-colour, 7 — 8 ju, diameter. Granules of lime very numerous, minute, hyaline, globose, or irregularly angular. Didymium Paraguaycnse, Spegazzini, Fung. Guarauit., n. 320. On fallen leaves and decayed wood. Guarapi. Didymium daedaleum, B. and Br. Sporangia connate, simcous, forming a dacdalioid mass, reddish- brown inclining to orange like the stem, sprinkled with white meal; stem connate, as if composed of a bundle of little flat membranes ; capillitium white, vei^y variable in undth, being in jjarts broad, fiat, membranous; spores violet-black, globose, smooth. Didymium dacdalcuvi, B. and Br., Ann. Nat. Hist., n. 885, p. 336 (1850); Cooke, Hdbk., n. 1131; Cooke, Myx. Brit., p. 35 ; Sacc, Syll., vii., 1, n. 1313. In a cucumber frame. Britain. There is no specimen of the present species in the Berkeley Herbarium, hence I am unable to add anything to the above somewhat imperfect description. Lejndodannfi. 251 Didymium connatum, Peck. Peridium depressed or subglobose, cinereous, furfuraceous, stipitate; stems mostly connate at the base, tapering upward, lono-itudinally wrinkled, whitish or cream-colour; spores sub- olobose, black, -OOO-i in. in diameter. Didymium cotmatiom, Peck, 26th Report, N. Y. State Mus., p. 74. Fhysarnm polymoiyhum, Sacc, Syll., vii., 1, n. 1196. Decaying fungi. Portville, U. States. The subfasciculate mode of growth is characteristic of this species. (Peck.) Unknown to me. In Saccardo's Sylloge the present species is oiven as a synonym of Physarwn polymorphum, but the description above does not justify this, and it is very doubtful whether the type specimen has been examined by Berlese. Didymium flavidum, Peck. Peridium subglobose, sessile, thin, yellow, clothed with a minute yellow mealiness ; spores subglobose, black, "0004' in diameter. Didymium flavidum, Peck, 28th Report, N. York State Mus., p. 54. Physarum Jlavidum, Berlese, Sacc, Syll., vii., 1, n. 1215. Bark of dead balsam trees, Abies halsamca. North Elba; U. States. Lepidoderma, De Bary, Sporangia sessile or stipitate or a plasmodiocarp, wall fur- nished externally with large scales consisting of bicarbonate of lime, superficial or enclosed in lenticular cavities of the mem- brane ; columella generally present. Rost., Mon., p. 187; Cooke, Myx. Brit., p. 43; Sacc, SyU., vol. vii., p. 386. Closely related to Didymium, and differing only in the fact that the external coating of lime is broken up into large, per- sistent patches, instead of forming a pruinose coating consisting 252 A Monograph of the Myxogastres. frequently of stellate crystals. Cliondriodcrma differs in having the external wall of lime forming a polished, porcelain-like crust. In the present genus the carbonate of lime is present in the form of very minute amorphous lumps until near to maturity, when it is dissolved and reappears as bicarbonate of lime deposited in comparatively large Hakes. The same change takes place in Physanoii granulatum, Balf. fil., and Mr. Harold Wingate, of Philadelphia, has remarked the same conversion of carbonate of lime into the bicarbonate in Chondriodcriaa Michdii, and adds, " I feel positive it takes place after tlie plant has matured, and possibly after it has been dry and again subjected to moisture." Distrih. Europe ; West Indies ; Ceylon ; Java, Species 8. A. Spores smooth. Lepidoderma stellatum, Mass. (figs. 223 — 225), Sporangia spherical, stipitate, slightly or not at all umbilicate below, UacJcish-hrown, variegated with rather large lohite sccdes, ruptured at maturity nearly to the base in a stellate manner, into 4 — 6 acute, irregular segments; stem rather thick, erect or rarely curved, striate, white; coluviella hemispherical or suh- clavate, rather prominent, whitish or tinged yellow ; threads of the capillitium very slender, branching at acute angles, hyaline, flexuous; spores smooth, dingy violet, globose, 10 — 12 fx diam. Grevillea, vol. xvii,, March (1889). On rotten wood. Dominica (Ramage). A very fine and distinct species, 2 5 — 3"5 mm. high ; scattered or gregarious. When young the sporangia are pure white, the outer coat becoming broken up into large white distinct scales as the sporangium expands. Readily recognized by the white stem, colourless capillitium, whitish columella and smooth spores ; the mode of dehiscence is also characteristic. Lepidoderma reticulatum, Mass. Sporangia gregarious, sessile on a hroad base, convex, circular or elongated, often Jlexuous and anastomosing in a reticulate Lejndodernia. 253 manner, wall thick, brown with a tinge of purple, furnished with large, scattered, innate, yellow patches of lime ; columella absent ; mass of spores purple-brown ; capillitium very scanty, threads 1 — 2 mm. thick, colourless ; spores globose, sniuoih, 11 — 12 fM diameter. Licea reticulata, B. and Br., Journ. Linn. Soc, vol. xiv., p. 86 (1873). Didymium reticulaium, B. and Br., in Herb. Berk. On bark and mosses. Ceylon. Distinguished from Lcjndoderma Chailletii by the absence of a columella, and from Z. Carestianum by the smooth spores and yellow patches of lime on the wall of the sporangium. Lepidoderma fulvum, Mass. (n. sp.) (figs. 39, 40). Sporangia stipitate, hemispherical, depressed, umbilicate hQueoXh., fulvous, with scattered, large, suiter ficial tvhite or yellowish scales ; stem stout, equal, or very slightly attenuated upwards, straight or a little bent, longitudinally rugulose, fulvous, hypo- thallus well developed ; columella large, hemispherical, brownish ; capillitium copious, threads slender, hranchcd at acute ' angles, flex%bou>s, Irownish-molct ; spores globose, smooth, brownish-purple, 10 — 12 IX diameter, some few much larger. On mosses, twigs, &c. Britain (Yorks.) ; France. (Type in Herb. Berk., 10,783.) In groups of 2 — 4, springing from a firm, well-developed hypothalluB, 2-5 — 3 mm. high. Differing from Zcjndoderma stellatum, the only other stipitate, smooth-spored species, in the coloured columella and stem. The calcareous scales on the pileus very thin, glistening, not so regular and distant as in L. fulvum. B. Spores vmrtcd. Lepidoderma tigrinum, Rest. (figs. 41 — 43). Sporangia hemispherical, depressed, stipitate, umbilicate beneath, Uachish-hrown, and furnished with rather distant, large yellowish scales ; stem erect, thick and equal, or slightly attcnu- 254 A Monograjih of the Mijxogastres. ated upwards, brownish rust colour or dingy ochre, wrinkled longitudinally ; columella hrown, globose or hemispherical ; threads of capillitium thin, brownish-violet, branches forming acute angles, sometimes united laterally ; spores globose, brown- ish-purple, minutely wartcd, 10 — 13 /x diani. Host., Mon., p. 187, figs. 159, 160; Cooke, Myx. Brit., p. 44, figs. 159, 160; Sacc, Syll., vii., p. 386. On mosses, wood, &c, Britain (Teesdale, Yorks. side ; New Pitsligo) ; France ; Germany ; Finland ; Sweden ; Ceylon. Superficially resembling Lepidoderma stcllata before dehi- scence, but known at once by the warted spores, which also separate the present species from L. fulva. Scattered or gregarious, 2 — 2'5 mm. high, the base of the stem expanding into a small, irregular hypothallus. Threads of capillitium sometimes with slight granular swellings. (Rostafinski's Synonyms.) Didymmm tigrinum, Schrad., t. 6, f. 2, 3 (1797) ; B. and Br., N. H., 383; Cke., Hdbk., n. 1121. Physarum tigrinum, Pers., Syn., 174 (1801); FL, Dan., t. 1434, f. 2. Physarum squamulosum, Pers., Syn., 174 (1801). Trichia tigrina, Poir., Ency., viii., 53. Tricliia squamidosa, Poir., Ency., viii., 53. Cionium tigrinum, Lk., Hdbk., iii., 410 (1833). Didymium riijijjes, Fr., S. M., iii., 116 (1829). Leangium squamulosum, Fr., Stirp. Femsj., 13 (1825). Lepidoderma obovatum, Mass. (n. sp.) (figs. 45 — 47). Broadly obovate, stipitate ; wall dirty ochraceous, thick and firm, studded with large, innate whitish patches of lime ; stem short, thick, dark hrown, wrinkled ; mass of spores blackish with purple tinge; columella absent; capillitium springing from the base of the sporangium, threads dingy violet, 3 — 4 /^ thick, equal, repeatedly branching in a dichotomous manner, furnished with a sioelling at the base of each dichotomy, the whole com- Lepidoderma . 255 billed into an irregular net; spores globose, dingy violet, minutely wartcd, 11 — 13 ju. diameter. On grass and twigs. Sweden. (Type in Herb., Kew.) Plant 1*5 mm. high, 1 mm. diameter; characterized by the obovate sporangium and the swelling at the base of each bifurcation of the capillitiura. The minute warts on the spores often show a tendency to become elongated. Lepidoderma Chailletii, Rost. Sporangia hemispherical, adnatc hy a hroad base to a hypo- thallus, blackish-violet, with numerous shining brown pro- tuberances; columella very small, brownish; threads of the capillitium subviolaceoics, loith scattered swollen poo^tions, forming a dense net ; spores subviolaceous, toarted, 10 — 12 ju diam. Rost., Mon., p. 189, fig. 179 ; Sacc, Syll., vii., p. 887 ; Cooke, Myx. Brit., fig. 179. France ; Germany. Lepidoderma Carestianum, Rost. PlasmodiocarjJ hroadly effused, pulvinate, flattened, wall mem- branaceous, fragile, greyish brown, with numerous rather small white squamules of lime ; threads of capillitium brownish gxey, yellowish, or almost colourless, branched, and sometimes slightly rough with minute granules of lime; spores globose, warted, brown with tinge of purple, 14 — 15 /u. diam. Rost., Mon., p. 188 ; Sacc, Syll., vii., p. 387. Beticularia Carcsiiana, Rabenh., MS., and in Rab., Fung. Eur., n. 436. On twigs. Italy ; Belgium. A peculiar species surrounding thin twigs like a cushion, 1 mm. thick, and extending for 1 cm. or more in length. Scetion douUful. Lepidoderma Kurzii, Berk. Sporangium globose, stipitate, nodding, obscure ycllov:ish- 256 A Monograph of the Myxogastres. green with siolphnr-coloured ixUchcs of lime ; stem slender, attenu- ated upioards and curved, sulphur-coloured, or whitish, becoming brownish towards the base ; hypothallus spreading ; spores blackish-brown, globose. Berk., in Herb., n. 10,785. On palm. Java. The above description, accompanied by sketches, is in Berke- ley's Herbarium, but the specimen has disappeared, hence I am unable to supplement the diagnosis. Spumaria, Pers. Aethalium consisting of numerous crowded, irregularly branched sporangia, the whole at first surrounded by a common friable cortex containing granules of lime ; sporangia dendritic, the thin walls containing granules of lime; columella central, not containing lime, sending off branches into the lobes of the sporangium; threads of the capillitiuin forming a dense network, extending from the columella to the walls of the sporangium. Sjmmaria, Pers., Disp., t. 1, figs, a, h, c; Rost., Mon., p. 191 ; Cooke, Myx. Brit., p. 45 ; Sacc, SylL, p. 838; Zopf, 153. The present genus, along with Diaehaca, form a transition from the Lithodermcae to the Gohimelliferae ; showing affinity with the former in the presence of granules of lime in the sporangial walls, and with the latter in the central, elongated columena. Distrih. Europe ; United States ; Canada ; Australia ; S. Africa; Ceylon. Species 1. Spumaria alba, D. C. Common cortex whitish, deciduous; sjwrangia ninnerous, dendritic ; columella empty, hranchcd, shorter than the branches of the sporangium, capillitium springing from the columella and passing to the wall of the sporangium, branches 2 — 4 ju thick, foi^ming a dense irregidar net, widened at the angles, obscure violet or brownish ; sjDorcs globose, mrructdose, obscure purple, 10 — 13 IX diameter. Spumaria alba, D. C, Flor. Fr., ii., 2G1 ; Rost., Mon., p. 191, Spumana. 257 figs. 158, 172, 175; Cke., Myx. Brit., p. 45, figs. 158, 172, 175; Schroeter, p. 120 ; Raunk., Myx. Dan., p. 89 ; Sacc, Syll, 1338. ^a;sicc.— Rab., Fung. Eur., 1323; Lib., PI. Crypt. Ard., Fasc. IL, 179 ; Karst., Fung. Fenn., 497 ; Fckl., Fung. Rheu., 1471 ; Syd., Myc. March., 397; Desm., Cr. Fr., ser. I., 768; Kunze, Fung. Sel., 198. On wood, bark, leaves, twigs, grass, &c. Britain (Kew, Scarboro', Carlisle, Capel Ciirig, Aboyne, N. B.) ; Germany ; France ; Belgium ; Italy ; Hungary ; Denmark ; United States ; Canada ; Australia ; S. Africa ; Ceylon. The Plasmodium often creeps up the stems of grasses for a distance of several inches, forming aethalia varying from I — 2 inches or more in length, at first covered with the thick, white, common cortex, which soon falls away, leaving the numerous, crowded, branched sporangia arranged in a dendritic manner, and of a bluish-grey colour. Mass of spores blackish, very copious and pulverulent. (Rostafinski's Synonyms.) Mucilago, 2, Mich., t. 96, f. 2 (1729). Mucorii, Gled. Meth., p. 160 (1753). Mucilago Crustacea alha, Batt., t. 40, f. 9, H. I. (1755). Mucilago, Hall, n. 2129 (1768). Byssus homhycina, Retz., V. Handl., 251 (1769). Mucilago filamentosa, Bonamy., t. 3 (1772). Beticularia alha, Bull., t. 326 (1791). Sjmmaria mucilago, Pers., Disp., t. 1, f. ct, h, c (1797). Beticularia ovata, Var., With. Arr., iv., 1978 (1803). Sjyumaria cm-nuta, Schum., Saell., 1415 (1803); Fl. Dan., t. 1978, f. 1. Spumaria alba, D. C, Fl. Fr., ii., 261 (1805); Fr., Syst. Myc, iii., 25; Eng. Fl., v., 310; Cooke, Hdbk., n. 1103. Spumaria alha, a. laminosa, (3. cornuta, Fr., S. M., iii., 95 (1829). Didymium spumarioidcs, Fr., S. M., iii., 95 (1829). Diderina spumariacformc, Wallr., Fl. Germ., 2208 (1833). 258 A Monograph of the Mi/xofjastres. DiACHAEA, Fr. Wall of sporangium without lime externally, but sometimes containing scattered granules on its inner surface, tliin, usually with metallic tints ; columella either thick and elongated, rigid with amorphous lumps of lime, or more or less rudimentary and represented by an accumulation of lime at base of spor- angium; capillitium forming a dense net, springing from the columella or base of sporangium, threads usually coloured, without lime, thickest at the point of origin. Diachaca, Fries, Syst. Orb. Veg., i., p. 143 ; Fr., Syst. Myc, iii., p. 155; Kost., Mon., p. 190; Cooke, Myx. Brit., p. 44; Sacc, Syll, p. 387. Although placed in a different order by Rostafinski, Lami^ro- dcrma approaches the present genus so closely, that it is open to doubt whether the two genera, as at present understood, should not be merged into one. The generic diagnoses of the two, as given by Rostafinski, illustrate the two poles of the genus in the larger sense, and read very distinct, but when a complete sequence of the species included in the two genera are examined, then the difficulty of " drawing the line " becomes only too apparent, and the only character that remains is, the presence of lime in the columella and stem iu Diachaca^ and its absence in Lam'proderma. In Laminodcrma violacca, Fr., the columella is filled with large vesicles as in Trichia fcdlax, and in Lampo'odcrma sulweneus, B., well-developed granules of lime are by no means rare in the threads of the capillitium ; lime is by no means always absent from the capillitium of several species of Stemonitis, and other genera belonging to the order Amaurochacte, the most important feature of which consists in the absence of lime. DiacJmea is as far from being a typical member of the Calcarcae, owing to the entire absence of lime on the surface of the wall of the sporangium, as it is from being ty])ical of the Amaurochaetcae on account of the lime contained in the columella and stem. Distrih. Europe; United States; W. Indies; Brazil; S.Africa; India; Australia; New Zealand. Species 5. Did elided. 259 Sub- Gen. DUirliaca. Columella well developed, elongated, rigid with amorphous masses of lime. A. S2)orcs smooth. Diachaea confusa, Mass. (n. sp.). Gregarious, springing from a thin, irregular, yellowish hypo- thallus ; sporangia elliptico-cylindrical, stipitate, obscure purple, iridescent, stem equal to, or a little shorter than the sporangium, 2Jale ochraccous, suhequal, slightly wrinkled longitudinally; columella two-thirds the height of the sporangium, (Hrty imh ochraccous, suhclavate, filled with masses of lime, very brittle ; threads of the capiJlitium springing from the columella, much branched and irregularly anastomosing to form a very dense net, about 3 /x thick near the columella, tapering, pale olive or smoky; s]Jores at first in clusters of 6 — 8, smooth, globose, 5 — 6 ^ diameter, dingy lilac. (Type in Herb., Kew.) Sporangia 2'5 — 3 mm. high ; distinguished by the ochraceous subequal stem and the small spores in clusters of 6 — 8. The capillitium is very dense, and the columella so very brittle that it requires care to see it intact. On living geranium leaves. Jamaica. Diachaea leucopoda, Rest. (figs. 165, 166). Gregarious, springing from a spreading white hypothallus ; sporangia elliptico-cylindrical, stipitate, brownish-purple, some- times iridescent, often subumbilicate below ; stem shorter than sporangium, white, thin above, cxjmnding eonsideraUy dmcmcards, smooth ; passing upwards as a thick, equal or slightly tapering coluviclla aJiout two-thirds the length of the sporangium, white, and filled with granules of lime ; mass of spores blackish ; capillitium dense, threads springing from the columella whore they are 3 — 4 fx thick, much branched and iiTegularly anasto- mosing to form a dense net, bccominq: thinner and attached 260 A Monograijli of the Mi/dcogastres. to the wall of the sporangium, dusky or almost colourless ; spores globose, minutely vcrnoculose, lilac, 7 — 9 fx diameter. Diachaea kucopoda, Rost., Mon., p. 190, f. 178; Cooke, Myx, Brit., p. 44, f. 178 ; Sacc, SylL, n. 1335. Exsicc. — Rab., Fung. Eur., 434 and 1071 ; Roum., Fung. Gall., 3174 ; Klotzsch (Rab.), Herb. Myc, 86 ; Syd., Myc. March., 1900 ; Rav., Fung. Car., 80 ; Desm., Cr. Fr., Fasc. ii., 261 ; Fckl, Fung. Rhen., 1451 ; Westendorp, Crypt. Belg., 1260. On leaves, twigs, &c. Britain (King's Cliff e, Carlisle, Lin- lithgow) ; France ; Germany ; Belgium ; United States ; Cuba ; Brazil ; S. Africa ; India ; E. Australia. Varying from 1'5 — 2 mm. high, characterized by the white, acutely conical stem, and smooth spores not produced in clusters. (Rostafinski's Synonyms.) Trichia kucopoda, Bull., t. 502, f. 2 (1791). Stcmonitis ekgans, Trent., in Roth., Cat., 220 (1797). Stcmonitis kucostyUi, Pers., Syn., 186 (1801). Stemonitis l&itcopoda, D. C, Fl. Fr., ii., 257 (1805). Diachaea ekgans, Fr., Stirp. Femsj., p. 84 (1825); Fr., S. M., iii., 156; Berk., Ann. N. H., n. 112; Cooke, Hdbk., n. 1154, f. 131. Diachaea kucopoda, Rost., Mon., p. 190 (1875). Var. cylindrica, Mass. Sporangium cylindrical, elongated, stem cylindrical. Exsicc— mXiB, N. Amer. Fung., 336. On pine wood lying on the ground. United States. Differs from the type form in its larger size, 2'5 — 3 mm. high; the longer cylindrical stem, and elongated, cylindrical sporangium. Diachaea Hookeri, Mass. Sporangia spherical, slightly flattened or umbilicate below, wall thin, pale ochraceoics or iviih a Jicsh tinge, slightly iridescent; with minute granules in detached patches on the inside ; stem Diarhaea. 261 about equcal to sporangium, stout, expanded at the base into a circular liypothallus, longitudinally rugulose, colour of spor- angium ; columella pale, cylindrical, ahmit half the height of the sjiorangiuiii, containing irregular masses of lime; threads of capillitium springing from the columella, irregularly branching and anastomosing to form a dense net, attached by its ultimate branches to the wall of the sporangium, threads 3 — 4 /x broad near the columella, flattened, tapering, pale dirty lilac to colourless; spores globose, minutely warted, brownish lilac, 11 — 14 }x. diameter. Didcrma Hool^eri, Berk., Fl., New Zealand, p. 191, T CV f. 12. Lamproderma Hookeri, Rost., Mon. Append., p. 24. (Type in Herb., Kew.) Gregarious on living fronds of Ilyvmiojjhylhm. New Zealand. A true Diachaea, as the genus is at present defined, and well marked by the pale coloured globose sporangium, and the large spores. (Rostafiuski's Synonym.) Diderma Hookeri, Berk., FL, New Zeal., p. 191, T. CV., f 12. Sub-Gen. Hemidiachaca. Columella short or obsolete, and represented by a few lime granules at base of sporangium. B. Spores ivith irregular, raised hands. Diachaea splendens, Peck (figs. 37, 38). Sporangia spherico-dq)rcsHcd and u/nhilicate heloiu, or rcniform, solitary or fasciculate on a common stem, deep peacock-blue or purple, iridescent ; stem equal to or shorter than sporangium, white, expanding into a vague liypothallus; columella white, cylindrical or reniform, containing lime; capillitium dense, threads brown, anastomosing to form a network extending from the columella to the sporangial Avail; spores globose, purple- 262 A Monograph of the Mi/,ro(/astres, brown, with scattered, broad, irrcgularly-shapcd raised hands of various lengths, 8 — 11 ju. diameter. Diachaea spkndcns. Peck, 30th Report, State Mus. Nat. Hist., p. 50; Sacc, Syll., 1336. On leaves, &c. United States. For specimens of the present very distinct and beautiful species, I am indebted to Mr. Harold Wingate, of Philadelpliia. Gregarious, about 1'5 mm. high, evidently allied to Diachaea kucojxida, but abundantly distinct in the spore marking, and the short reniform sporangia and columella. Diachaea suhsessilis, Peck. Gregai'ious or crowded ; sporangia subglobose, sessile, or with a very short white stem, the walls delicate, iridescent with various metallic tints; columella ohsolete ; capillitium and mass of spores violet-brown ; spores globose, rough, '0004' — 'OOOo' in diameter ( =■ about 12-5 fx ). Diachaea sicbsessilis, Peck, 31st Report, State Mus., p. 41. Fallen leaves. Adirondack Mountains, U. States. This is a most singular species, and apparently very rare. In its lack of a distinct columella, it departs from the generic character, but it cannot be placed in the genus Lamproderma, for the same objection would hold there, and besides that, another is found in the presence of lime granules in the stem. Even when no distinct stem is present, a small whitish mass of granules can generally be seen at the point of attachment. The capillitium appears to originate at the base of the spor- angium. The spores are larger, but less rough than those of D. splendcns. (Peck.) I have had no opportunity of examining the present species, hence can add nothing to tlie vague description of the epispore. Subord. Physarae. Craterium, Trent. Sporangia usually distinct and stipitate, wall consisting of two or three distinct layers, dehiscing by a sharply-differentiated Crateriiiin. 263 apical lid, or by an irregularly circuinscissile separation of the undifferentiated apical portion of the sporangium, basal portion of sporangium persistent after dehiscence ; capillitium of stout threads attached at numerous points to the sporangial wall, combined to form an irregular, rigid network, containing a large quantity of lime which is usually localized in swollen, irregular, often more or less flattened portions ; these towards the base of the sporangium are compacted to form an irregular columella. Crateriuvb, Trent., Obs. Bot. ; Rost., Mon., p. 118; Cke., Myx. Brit., p. 18 ; Zopf, p. 146 ; Sacc, Syll, vii., 1, p. 355. The species of Craterium are more or less gregarious in habit, but rarely crowded, although now and again specimens occur with the sporangia more or less confluent, the stems remaining distinct. The sporangia persist for a long time after dehiscence, owing to the rigidity of the wall and capillitium, and in this stage resemble in shape miniature wine-glasses. Craterium is closely allied to BadJiamia, difl'ering more especially in the double wall and the presence of a columella, and the circum- scissile dehiscence. Distrib. Europe ; Algeria ; Natal ; Ceylon ; Tasmania ; Brazil ; United States. Species 13. A. Dehiscence hy the sejmratiou of a disthict lid at apex of S])oi'angium. Craterium confusum, Mass. (fig. 280). Sporangia variable in form, cyathiform, infundibuliform, or pyrifonn, stipitate or rarely subsessile, varying in colour from bright brown, through ochraceous to nearly white, lid sharply differ entiafcd from wall of sporanijinui, flat or slightly convex, ■HSU((Ily chalky-white, rarely ochraceous, deciduous ; stem variable in length and depth of colour ; capillitium abundant, knots of lime large, irregular, white or yellowish, usually by aggregation forming a more or less distinct columella ; spores globose, dirty violet, vei'y minutely verrucnlose, 8 — 11 ii diameter. Craterium vulgare, Rost., Mon., p. 118, figs. 94, 9G ; Cke., Brit. Myx., p. 18, figs. 04, 0(J. 264 A Monograph of the Myxogastres. Craterium i^yriformc, Rost., Mod., p. 120; Cke., Brit. Myx., p. 19; Sacc, Syll., n. 1234. Cratcrium minntum, Rost., Mon., p. 120, f. 95; Cke., Myx. Brit, p. 19, f. 95 ; Raunk., Myx. Dan., p. 79. Craternim 2'>cdunculatum., Sacc, Syll., 1233; Raunk., Myx. Dan., p. 79. Exsicc.—QkQ., Fung. Brit, Ed. II., 208 (as G. minutum); Fckl., Fung. Rhen., 1453; Roum., Fung. Gall., 2957; Syd., Myc. March., 489; Desm., Cr. Fr., Ser. I, 869; Fckl, Fung. Rhen., 1454 (as Cratcrium pyriforme). On leaves, twigs, bark, moss, &c. Britain (Kew, Weybridge, Penzance, Apethorpe, Bungay, Lyme Regis, Shere, Scarboro', Carlisle, Abergavenny, Appin, Glamis, N. B.); Germany; Hungary; France; Italy; Finland; Belgium; Bavaria; Sweden; United States ; Cuba ; Brazil ; N. Zealand ; Australia ; Natal ; Tasmania. After a careful examination of numerous specimens from Europe and other parts of the world, I find that the only course left is to combine the three species and five varieties acknow- ledged by Saccardo, into one species characterized by the sharply- differentiated operculum, which is usually chalk-white, and flat or slightly convex, and the dull-coloured sporangium. In the above idea I am confirmed, or rather anticipated, by Mr. A. Lister, who, during a critical examination of the Myxogastres in the Kew Herbarium, has attached the following note to the specimen-sheet of G. vulgarc. " I have added to the col- lection some specimens of Graterium viUgare, representing a few varieties in form and colour from a heap of dead leaves in my grounds at Lyme Regis. The white form is abundant in such localities, and given in several of the Kew specimens, is the result of age and exposure, on originally more deeply coloured, but pale sporangia. I have cultivated a large number of sporangia from orange-yellow plasmodium, and gathered many thousands of the ripe fruits during the last three years from this particular heap. Among them I find all the characters given in the description of Rostafinski's three species, G. vidgarc, G. inirifornu, and G. rninuiuin, with (as it Cratermm. 2G5 appears to me) all shades of intermediate form. From careful observation of sporangia developed from plasmodium, botli in the open air and mider bell-jars, I believe that while varying in form and colour, they belong specifically to one species. The Plasmodium appears to be always ultimately, of a more or less orange-yellow, though in some stages it is often dirty brown from the quantity of foreign matter held in suspension. The differences in form and colour of the sporangium are the only characters by which these three species (?) are separated, and these characters vary so much in sporangia rising simul- taneously, and apparently from the same plasmodium, that, I venture to submit, they cannot be considered as specific. " In all varieties the wall of the cup consists of two or three layers, the outer smooth, with granules of lime equally dis- tributed throughout its more or less coloured substance, the inner white, and heavily charged with lime, though the lime differs in amount even in sporangia of the same cluster. The lid is deciduous, and usually paler than the cup, and in most instances distinctly marked off from the rim. In abnormal forms, caused by rapid drying, the lid is often sunk below the rim. " The capillitium is white, consisting of large knots of lime connected by scanty hyaline threads; these threads are some- times yellow, when the colour extends over the inner wall. A central aggregation of lime knots or columella is sometimes present. Spores smooth, 5 — 9 /x, bright violet or violet-brown." The appearance of the epispore depends on the amount of magnification. I find in Mr. Lister's own specimens that under a power magnifying 1200 diameters the epispore is distinctly warted. (Rostafinski's Synonyms.) (Under C. vulgare.) Fangoides wfumlihiliforine, pendulum, pediculo donatum, Mich., Nov. PL Gen., p. 205, n. 10, t. 86, f 13 (1729). Crateriiim 2Jed%onculatitm., Trent., I.e., p. 2-i-i (1797). Cratermm mdgarc, Ditm., I.e., t. 9; Nees, Syst., f. 120; Chev., Fl. Par., t. 4, f. 2G (1817). 266 A Monograph of the Myjcogastres. Cratcrium hucocephahtm , Desm., Cat. 27, non Auct. (1823). (Under C. 2)yriforme.) Craterkwi 2)1/^/0^116, Ditm., I.e., p. 19, t. 10 (1817). (Under C. viinutum.) Peziza minuta, Leers., Fl, Herbor., n. 1085 (1775). Cyathus minutus, Hoff., Veg. Crypt., p. 6, t. 2, f. 2 ; Sow., Eng. Fg., t. 239 (1790). Trichia minuta, Relh., Teste Fr. (1786). Nidularia onimcta, With. (1792). Sphaerocarpa operculata, Scliuni., Fl. Saell., n. 1503 (1803). Physarurii turbinatum, Schum., Fl. Saell., n. 1450 (1803). Craterium leucocephalum, Grev., Fl. Scot., t. 65 (?) (1824). Craterium minutum, Fr., I.e., iii., p. 151 (1829). Ci'atermm turbinatum, Fr., I.e., iii., p. 152 (1829). Arcyria leucoce])liala, Auct. Craterium Oerstedtii, Rest. Sporangia pyriform, along with the stem hlacJcish-lroim, lid convex, snotv-v;Jdte ; columella distinct ; spores pale violet, 8 — 10 fl diameter, smooth. Craterium Oerstedtii, Host., Mon., p. 120, fig. 99; Cke., Myx. Brit., fig. 99 ; Sacc, Syll., vii., 1, n. 1235. On wood. Denmark. Unknown to me. Appears to differ from C. confusum in the darker-coloured stem and sporangium. Craterium Friesii, Rost. (fig. 265). Sporangia cyathiform, stipitate, wall smooth, shining, orange- red, lid distinct from cui^, lohitish, flattened, stem very thin, equal, longer than sporangium, red; capillitium dense, knots of lime large, joined by short, hyaline internodes ; spores globose, brownish-violet, smooth, 10 — 11 ix diameter. Craterium Friesii, Rost., Mon., p. 122, f. 105; Cooke, Brit. Myx., f. 105. Cratcrium nutans, Sacc, Syll, 1237. Craterium. 267 On mosses, decaying lichens, &c. Britain (Lyndhurst, Haw- thornden, Edinboro') ; Sweden ; Germany. From 1"5 — 2 mm. liigli, sporangia and stem varying from orange to deep blood-red, stem elongated, slender. B. A'pcx of siwrangium not differentiated into a distinct lid. Craterium leucocephalum, Rost. (figs. 298 — 301). Sporangia stipitate, conico-cylindrical, pyriform or subglobose, apex rounded and eventually Ireaking away in a circumscissile manner, upper portion of sporangium whitish and rough loith minute, inmate particles of lime which eventually fall away, leaving minute pits in the wall, basal portion of sporangium dark reddish-brown, much wrinUed ; capillitiura dense, knots large, irregular, numerous, usually aggregated in the centre to form a columella, granules of lime colourless, yellow or brownish ; spores globose, dull violet or brownish, mimUely ivartcd, the wart often with a tendency to form anastomosing lines, 8 — 11 i^ diameter. Craterium leueocei^halum, Eost., Mon., p. 123, f. 98, 100; Cooke, Myx. Brit., p. 19, f. 98, 100 ; Raunk., Myx. Dan., p. 80, t. 4, f. 12; Sacc, Syll., 1238. Exsicc.—'^ydi., Myc. March., 1500; Fuckel, Fung. Rhen., 1452; Roum., Fung. Gall, 1679; Rab.-Winter, Fung. Eur., 2G74. On leaves, twigs, bark, &c. Britain (Jedburgh, Chiselhurst, Epping, Gloucester, Wanstead Park, Scarboro', Carlisle, Appin, N. B.) ; Germany ; France ; Italy ; Denmark ; United States ; Brazil. PJasraodium yellow. Gregarious or scattered, 1'5 — 2 mm. high, variable in form and length of stem, the upper convex portion of the sporangium breaks away in a circumscissile manner, there is not a differentiated lid as in C. vulgarc. The present species is best characterized by the pale upper portion of the sporangium being incrusted with minute particles of white or yellowish lime, and also by the presence of numerous circular discs sunk in the substance of the wall; these discs vary 268 A Monograph of the Myxogastreif. in size from 10 — 50 [i diameter, have the margin more or less grooved, and show indications of hnes radiating from the centre to the circumference. Tliese discs occur also rarely in the capillitium threads. (Rostafinski's Synonyms.) Fungoides minimum infundibiliforme, albicans, pediculo dona- tum. Mich., Nov. PL Gen., p. 205, n. 13, t. 86, f 14 (1729). Peziza convivalis, Batsch., Elenoh. Fung., p. 121 (1781) (not convivalvis, as erroneously quoted by Rostafinski and Cooke). Stemonitis leucocephala, Pers., in Gmel., Syst, Nat., p. 1467, n. 9 (1791). Stemonitis cyathiformis, Schrnk., I.e., p. 19 (1790). Trichia cinerea, Trent., I.e., p. 227 (1797). Arcyria ImLCocephala, Hoffra., Fl. Or. ger., t. 6, f 1 (1795). Physarum pcdunmlatum,, Schum., Fl. Saell., n. 1453 (1803). Trichicce aureae affinis, Fl. Dan., t. 1314, f. 2 (1810). Cyathus cinereus, Purt., Brit, iii., n. 1561, t. 35 (1817). Crateriwn leucocepJicdum, Ditm., I.e., t. 11, Grev., Fl, Scot., t. 65 (1817). Craterium viilgare, Chev., Fl. Paris, 340, non Ditm. (1826). Physamm leucostictum, Chev., Fl. Paris, p. 336, t. 9, f 29 (1826). Craterium leucostictum, Fr., I.e., iii., p. 152 (1829). Gupularia hucocephala, Lk., Hdbk., iii., 421 (1833). Physarum xanthopus, Wallr., in Sched. (1833). Craterium xantJiojJUS, Wallr., Fl. Germ., ii., 358 (1836). Craterium deojoerculattim, Fr., in Wein., I.e., p. 597 (1836). Cupularia xanthopus, Rabh., Fl. Cr. Germ., n. 2226 (1844). Craterium pruinosum., Corda, Icon., vi., p. 13, t. 2, f. 33 (1854). Craterium cylindricum, Mass. (n. sp.) (fig. 213). Scattered or gregarious, stipitate, sporangium cylindrical, elongated, apex rounded, smooth, even, lower portion pale clear red, white above, upper portion breaking away in a circum- Crateriam. 269 scissile manner; stem thin, elongated, red, like base of spor- angium; capillitium dense, nodes very large, irregularly angular, filled with yellowish granules of lime, connected at various points by long, very thin internodcs not containing lime, com- bined at the base of the sporangium to form a short columella ; spores dirty lilac, globose, smooth, 9 — 10 fx diameter. Uxsicc.—EWis, North Amer. Fung., exs., n. 1400 (as Craterium leiicocephnlum). On twigs, &c. United States. Stem about two-thirds the length of the sporangium, 2—2-5 mm. high. Allied to C. leucoccjyhahim, but distinguished by the elongated, narrowly cylindrical sporangium, the long, thin internodes of the capillitium, and the smooth spores. Craterium aureum, Rest, (figs, 257 — 261). Sporangia elliptic-oblong, subglobose, or pyriform, stipitate, wall i\nck,hvittle,riigtdose,from hright lemmi-yellow to reddish- orange, upper convex portion breaking away in an irregularly circumscissile manner; stem usually shorter than sporangium, arange-yelloio, subequal, expanding at the base into a small hypothallus; capillitium dense with numerous large, irregular knots with yellow or white granules of lime, connected hy elon- gated, thin, emjJty internodes; knots aggregated in the centre towards the base to form a columella ; spores globose, brownish- lilac, miniLtcly verruculose, 8 — 10 [jl diameter. Craterium aureum, Rost., Mon., p. 124; Cooke, Brit. Myx., p. 20 ; Sacc, Syll., 1239. Craterium mutaUlc, Fr., S. M., iii., p. 154. On leaves, bark, twigs, moss, &c. Britain (Kcw, Carlisle, Lyndhurst, Appin, N. B.) ; Sweden; Germany; Algeria; United States. Plasmodium clear lemon-yellow. Scattered, 1-5 mm. high, distinguished by the rugulose, bright yellow, rigid sporangium, and the long slender internodes in the capillitium, in which respect the present species approaches the genus rhysarnm. 270 A Monograph of the Myxogastres. (Rostafinski's Synonyms.) TrkUa aurea, Schiim., Fl. SaelL, 1461 (1803). Craterium mutahik, Fr., I.e., iii., p. lo-i, non S. Gast. (1829). Cuimlaria mutalilis, Rab., Fl. Germ., Cr., n. 2225 (1844). Physarum Duricui, M,, Herb. Craterium rubiginosum, Mass. (figs. 309, 310). Sporangia globose, broadly elliptical, or obovate, stipitate, dehiscing in a more or less regular circumscissile manner, lower portion of sporangium permanent, reddisli-brown, often with a purple tinge, lid or apical portion paler, sometimes whitish ; stem about equal in length to sporangium, stout, dark red- brown, passing downwards into a spreading hypothallus; capillitium dense, white, threads aggregated in the centre to form a darh-coloured columella ; spores dingy violet, globose, minutely warted, warts sometimes more or less arranged in lines, 10 — 13 /^ diameter, Badhamia ruMginosa, Rest., Mon. App., p. 29 ; Sacc, Syll., vii., n. 1158. Scyphium riddginosum, Rost., Mon., p. 148, fig. 115 ; Cke., Myx. Brit., p. 29, fig. 115. Hxsicc.—EiMs, N. Amer. Fung., 1215. On wood. Britain (Appm, N. B.) ; Germany ; France ; United States. Gregarious, 2'5 — 3 mm. high. (Rostafinski's Synonym.) Physancm ruUginositm, Chev., Fl. Par., 338 (1826); Eng. FL, v., 315; Cke., Hdbk., n. 1137. Craterium dictyospermum, Mass. (figs. 805, 306). Sporangia globose or obovate, dehiscing in a chxumscissih manner or irregularly, lower portion of sporangium permanent, reddish-brown, becoming paler or whitish at the apex; stem about equal in length to the sporangium, rather stout, dark Craterium. 271 brown ; capillitiura dense, white, threads aggregated in the centre to form a hroumish columella; spores violet with usually a brown tinge, globose, covered with irregular simple or hranclicd ridges, ivhich here and there combine to form an imx)erfect network, 10 — 14 i*. diameter, Badhamia dictyos2wra, Rost., Mon. App., p. 4; Cke., Myx. Brit., p. 82. On wood, moss, &c, Britain (Carlisle) ; United States. Gregarious, 2 — 3 mm. high ; indistinguishable from C. ru- higinosuiii, except by the nature of the markings on the epispore. May possibly prove to be a form of the last-named species. (Rostafinski's Synonyms.) Physarum canum, Klotzsch., Hb. Houk. Physaritm rubiginosum, Berk., Eng. FL, v., p. 31; Craterium lilacinum, Mass. (figs. 307, 808). Sessile, aggregated, globose, smooth, reddish-lilac, capillitium threads variable in diameter, clustered together in the centre and forming an irregular columella ; spores purple, sometimes tinged with brown, globose, epispore furnished with short, irregular ridges, which sometimes combine to form an irregular network, 12 — 16 ju diameter. Badhamia lilacina, Rost, Mon., p. 145, figs. 108, 109; Cke., Myx. Brit., p. 27, figs. 108, 109 ; Sacc, Syll., vii., n. 1156. Exsicc. — Syd., Myc. March., 1297 (as Leocarims irrnicostis). On rotten wood, bark, twigs, &c. Britain (Carlisle); Ger- many ; United States. Usually crowded, sporangia seated on a broad base, wall generally smooth, but sometimes rather rough. The present species may eventually prove to be a sessile form of C. dietyo- spcrmum. (Rostafinski's Synonym.) Physarum lilacinum, Fr., I.e., iii., p. 141 (1829). 272 A Monograph of the Myxogastres. Craterium Curtisii, Mass. Gregarious, sessile, obovate, fixed by a point to a very thin hypothallus, umher-hroivn, wall tliin, smooth or sometimes rugu- lose, soon disappearing at the apex ; capillitium dense, nodes large, irregular, sepao^ated only hy short constrictions, everywhere containing pale brown granules of lime, the nodes become confluent towards the base and form an irregular columella ; spores brown with indistinct lilac tinge, globose, minutely warted, 7 — 8 ju, diameter. Badhamia Curtisii, Rost., Mon. App,, p. 3; Sacc, Syll., vii., 1, n. 1159. Didymium Curtisii, Berk., Grev., vol. ii., p. 65. (Type in Herb. Berk., n. 10,758.) " Bcypliium Curtisii, Rest., Mon., p. 149. On leaves, grass, &c. United States. Sporangial wall very thin, shining, containing very little or no lime. About 1 mm. high, Craterium Fuckelii, Mass. (n. sp.). Sporangia perfectly globose, contracted below into a short, stout stem, every part hrownish-ochre, upper portion breaking away in an irregularly circumscissile manner; nodes of capil- litium large, angular, combining towards the base to form an irregular columella; spores dirty lilac, globose, rather coarsely warted, 12 — 14 fx diameter. Craterium mutcdiile, Fr., in Fuckel's Fung. Rhen., Exs., n. 1455. (Kew copy.) On wood. Germany. Distinct from C. auretim in the globose, differently-coloured sporangia, the absence of the long, thin internodes of the capillitium, and the larger, coarsely-warted spores. Craterium minimum, B. and C. Sporangia broadly elliptical, apex convex, breaking away in an irregularly circumscissile manner, pale yellow, becoming Pln/mrum. 273 brownish towards the base, as is also the very short, equal stem ; nodes rather scanty, large, irregularly angular, connected at vririous 2ioints hy rather long, thin internodcs ; spores globose, dirty lilac, 7 — 8 ix diameter, smooth. Craterkim minimum, B. and C, Grevillea, vol. ii., p. 67 ; Host., Mon., p. 125 ; Sacc, Syll, vii., 1, n. 1240. (Type in Herb. Berk., Kew, n. 10,827.) On herbaceous stems, &c. Lower Carolina. Usually growing in lines on slender, dead herbaceous stems, •5 up to nearly 1 mm. high ; the long, thin internodes of the capillitium are unusual in the genus. Craterium minutum, Kickx. Sporangium stipitate, campanulate, rounded and umbilicate below, 1 — 1-5 mm. long, of a yellowish clay-colour, tinged brownish or reddish towards the base, sprinkled with very minute, wart-like, paler spots. Operculum the same colour as the upper part of the sporangium. Filaments whitish. Spores globose, black. Stem 2, rarely 3 mm. high, attenuated towards the apex, wrinkled, often compressed, russet-brown or reddish, springing from a blackish-brown indistinct hypothallus. Craterium. minutum., Kickx, Flor. Flanders, ii., p. 23. On rotten wood. The present species is considered as a species of Tilmadoche by Rostafinski, but as his description differs very materially from that of Kickx, the latter is given above. There is no reason, judging from the description alone, to show why Kickx was not quite correct in considering his specimen as a Crateriiom. Physarum, Pers. Sporangia solitary, plasmodiocarp, or combined to form an aethalium, wall single or of two distinct layers; threads of capillitium springing from all parts of the sporangial wall, combined to form an irregular network, with the numerous nodes or angles very much and irregularly swollen and filled 274 A Monorfraph of the My^rogastres. with granules of lime, the interuodes thin and without lime ; dehiscence irregular, or by a longitudinal fissure in the plas- modiocarp form. Physarum, Pers. Rost., Mon., p. 93; Cke., Brit. Myx., p. 11; Sacc, Syll., vii., p. 38G ; Zopf, p. 144. The principal characteristic of the present genus is the presence of numerous large, irregularly swollen nodes, or points of junction of capillitium threads, filled with granules of lime, the in te modes, or threads connecting the nodes, being thin and free from lime. Physarum passes by numerous transitional forms into Tilmadoche, which in the typical condition differs in the much fewer and smaller lime-containing nodes of the capillitium. Distrib. Europe ; Africa ; Himalayas ; Ceylon ; Java ; Japan ; W. Australia ; N. Zealand ; Tasmania ; S. America ; W. Indies ; TJ. States. Species, 70. A. Sporangia stvpitate. (Some species included in the present section have sessile forms.) * Ujrisjjore loarted. Physarum psittacinum, Ditm. (figs. 290, 291). Scattered, gregarious, or fasciculate, sporangia stipitate, globose, slightly rugulose, lime scanty, gree^i, often with orange and deep violet tints ; stem elongated, subequal or attenuated upwards, expanding at the base into a small hypothallus, longi- tudinally rugulose, orangc-rcd ; capillitium abundant, consisting of irregularly anastomosing, rather thin threads, connecting numerous large irregular nodes filled with lime, cither colourless or usually some shade of orange; columella absent; spores globose, hrawnish-purplc, very minutely verructdose, 7 — 9 /x diameter. Physarum j^sittaeinum., Ditm., Sturm, t. 62 ; Rost., Mon., p. 104, figs. 75, 76 ; Cooke, Brit. Myx., figs. 75, 76 ; Sacc, Syll., n. 1179 (Excl. Syn. Didymium, Ravenslii, B. and C.) ; Raunk., Myx. Dan., p. 74. Pliysartim. 275 Lidyminm crythrinum, Cke., Grev., 1873. On bark, &c. Britain (Neatishcad, Norths); Germany; Sweden ; France ; Italy ; Denmark. Distinguished amongst the red-stemmed species by the green sporangium, and the capillitiura consisting of thin, elongated internodes, some of the nodes are flattened, with one or more perforations and contain no lime, numerous others are filled with lime which is usually tinged orange ; from 2 — 2'5 mm. high. Care must be taken not to confound the present species with the green forms of TUmadocfie uiatahilis. (Rostafinski's Synonym.) Phi/sarum 2)sittaci)iwn, Ditm., I.e., t. G2 (1817). Physarum Schumacheri, Rost. (figs. 287, 288). Scattered or gregarious; sporangia globose, stipitate, warted with granules of lime, ycllmvish-olive, often with a tinge of green; stem erect, subequal, usually longer than the spor- angium, filled with lime, longitudinally rugulose, expanding nt the base into a small hypothallus, j)«/c yellow; columella cylindrical, whitish ; capillitium dense, forming an irregular net, with numerous rather smrdl knots containing yclloivish granules of lime ; spores globose, dull violet, minutely verruculose, 7 — 9 /i diameter. Physarwiih Schumachcri, Rost., Mon., 99 ; Cke., Myx. Brit,, p. 11 ; Raunk., Myx. Dan., p. 74. Physarum citrinum, Sacc, Syll., n. 1176. Physarum chrysojpeplum, B. and G., in Herb. Exsicc.—EWi?,, N. Amer. Fung., n. 1395. On bark, leaves, &c. Britain (Neatishead, Carlisle); Ger- many ; Italy ; Finland ; Ceylon ; United States ; S. America. About 2 mm. high. Capillitium dense, rather flaccid, inter- nodes long, thin, many of the angles flattened and not contain- ing lime, others irregularly branched and tilled with yellow granules; columella prominent. The above description applies 276 A Monograpli of the Myocogastres. to the typical form, but Rostafinski has given as forms of this species a heterogeneous collection, varying considerably in habit, . colour, and size of spores, but all agreeing in having a colu- mella; how many of these forms are British I do not know, /3. chryscypus, which appears to be the same as Physarum citri- 7inm, Schum., has been collected in Yorkshire, and a specimen probably corresponding to y. anrantiacum has been met with at Neatishead. The following are Rostafinski's forms. Var. a. genuinum. Sporangium yellow or greenish-yellow, 5 mm. wide, stem yellow ; spores 7 — 8 \x diameter ; granules of lime yellow. This variety is the one described above as the type form. /3. chrysojms. Sporangium 1 mm. diameter, with the stem golden-yellow; spores 10 [x diameter; granules of lime golden- yellow. y. cmrantiacum. Sporangia I mm. diameter ; stem brown ; spores orange, 11 /x diameter; granules of lime brown. 8. rufipes. Sporangium yellow or orange, sometimes iridescent ; stem orange-red ; granules of lime yellowish. e. compadum. Plasmodiocarp veinlike, creeping, short, yellow or golden ; columella absent, disappearing with the stem. Didymium melleum, B. and C, given as a synonym of the present species by Bostafinski, although a Physarum, is quite distinct, as shown by an examination of the type specimen in the Berkeley collection in the Kew Herbarium. (Rostafinski's Synonyms.) Physarum citrimtm, Schum., Saell., i., 436 (1803). Physarum aurantiacum, fi. ritfipes, A. and S., Cons., 262 (1805). Physarum verrucosum, Link, Herb. Physarum compadum, Ehr., Syl. Ber., p. 21 (1818). Physarum Schumacheri, Spr., Syst., iv., p. 528 (1sii, but differing in the structure of the capillitium, the nodes containing lime are much smaller, the internodes long and slender, and many of the nodes are empty and collapsed, or contain air only and not lime. Physarum candidum, Rost. Sporangia strictly or irregularly globose, sessile or stipitate, dehiscing irregularly, wall charged with lime, snoiv-tvhite ; stem luhite, elongated, plicate, 1 mm. high, rigid, crustaceous, capillitium with snoio-white roundish nodes containing lime; spores pale violet, murieatc, 12 — 15 fx diameter. Physarum eandidtcm, Rost., Mon., p. 9G ; Sacc, Syll., vii., 1, n. 1170. On rotten wood. Juan Fernandez. Physarum simile, Rost. Gregarious, stipitate, sporangia globose, gory, densely covered with small white lumps of lime, rigid ; stem usually twice as long as sporangium, equal, or slightly thickened below, and passing into a small, orbicular hypothallus, ycllmoish, sometimes with a pink tinge, rigid, filled with particles of lime, usually longitudinally wrinkled ; columella subcylindrical, whitish, one- fourth the height of the sporangium or less; capillitium dense, threads thin, colourless, forming an irregular network, stvcllings of nodes or interstitial parts sonall, elliptical or triangular, with colourless or yelloioish granules of lime; spores dingy lilac, very miiiutcd vcronic'idose, 7 — 8 fx diameter. Plijl^iwum. 287 Physarum simile, Rost., Mon. Append, p. 6 (species founded by Rostafinski on a specimen in Heib. Berk.); Sacc, Syll., vii., 1, n, 1175. Physarum Petcrsii, var. Farlowii, Rost., Mon. Append., p. 6 (specimen named by Rostafinski in Herb. Berk.). Krsicc. — Ellis, N. Amer. Fung., n. 1120. On wood, moss, &c. United States. Closely resembling P. Petcrsii, differing in the grey sporangium. About 1.5 mm. high. Physarum leucopus, Rost. (figs. 60 — 02). Sporangia globose, broadly ellipsoid or a little depressed, stipitate or sessile, rarely elongated and flexuous or anasto- mosing, wall at first covered tvith a continuous, snow-white coat of lime, which soon becomes hrohen up into smooth, innate 2Mtches ; stem variable in length, white, containing lime, straight, brittle, slightly thinner upwards, longitudinally rugose, passing into a more or less evident hypothallus ; columella absent ; capillitium strongly devclojjcd, snow-white, loith numeroiLS large, irregularly branched knots, containing lime in small granules; spores globose, dingy lilac, rather coarsely ivarted, warts almost black, 9 — 12 \j. diameter. /. stipitatum. Stem present, length variable, in the typical form, equal to, or longer than, the sporangium. /. sessile. Stem very short or entirely absent, when the sporangia are sessile on a broad base, sometimes confluent, elongated, sinuous, or anastomosing irregularly. Physarum leucopus, Rost., Mon., p. 101 ; Cke., Myx, Brit., 12; Schroeter, p. 129; Karsten, Myc. Fen., p. 102; Sacc, Syll., vol. vii., pt. I., n. 1188. Physarum Imcophacum., y. flc.cuosum, Cooke, Myx. Brit., p. 15. Exsicc. — Rab., Fung. Eur., 835 (as Pltysarum alhipcs); Cooke, Fung. Brit., Ed. II., 519. On wood, moss, &c. Britain (Highgate, Carlisle); Germany; Finland ; Sweden ; S. Africa ; S. America ; Ceylon ; Australia. A somewhat rare species, often represented in Herbaria by 288 A Monogra2'>h of the Myxogastres. Didymium sqi/amnlosum, which differs in the large, white columella. Distinguished from sessile forms of Fhysarum leu- copliaeum, by the coarser capillitium furnished with numerous large nodes, and the slightly larger and more coarsely warted spores. (Rostafinski's Synonyms.) Didymium Icucopiis, Link, Diss., 2, p. 42 (1809). Fhysaimm hullatum, Link, Diss., 2, p. 42 (1809); Ditmar, t. 22. Physaritm cdhopunctatum, Link, Herb. JDidymiiLm leucojms, Fr., S. M., iii., 121 (1829); Engl. FL, v., p. 313; Cke., Hdbk., n. 1127. Fhysarttm o-amentaceum, Fr., in litt. ad Wein (1836). Physarum leucophaeum, Fr. (figs. 63 — 66 and 96). Sporangia subglobose, usually a little depressed below, stipitate or sessile, wall thin, with irregular, white, innate patches of lime, dehiscing irregularly ; stem usually longer than the spor- angium, erect, slightly attenuated upwards, longitudinally rugu- lose, brown, paler above, passing into a dark hypothallus ; capil- litium dense, colourless, anastomosing very irregularly, threads thill, often flattened at the angles ivhich rarely contain lime; spores globose, dingy lilac, minutely warted, 8 — 10 fx diameter. /. fasciculatum. Stems confluent. /. sessile. Stem very short or entirely absent, sporangia scattered or confluent, regular or sometimes elongated and anastomosing. Physarum lcicco2)haeicm, Fr., Symb, Gast., p. 24; Rost., Mon., p. 113, figs. 77, 78, and 89; Cooke, Myx. Brit., p. 15, figs. 77 78, and 89; Sacc, Syll., n. 1192; Schroeter, 129; Raunk., Myx. Dan., p. 75. Didymium pruinosum, B. and C, Cuban Fungi, n. 530. (Type in Herb. Berk.) On leaves, twigs, bark, moss, &c. Britain (Queen's Cottage grounds, Kew ; Hampstead, Bristol, Deal, Scarboro', Carlisle, PItymriun. 289 Edinburgh); France; Germany; Italy; Hungary; Denmark; S. Africa ; Australia. About 1*5 mm. high ; stem, when perfect, clingy brown, slightly attenuated upwards ; the sessile and irregular forms are dis- tinguished from Physarmn cinereum by the dense, irregular capillitium having most of the angles flattened and rarely containing lime. Va7\ violasccns, Rost. Sporangium subglobose or compressed, wall thin, iridescent, with violet or reddish tints, with very few small, innate patches of lime, or these may be entirely absent; stem short or equal to sporangium, stout, expanding at the base into a small hypothallus, yellowish, darker below, strongly wrinkled longitudinally ; capillitium and spores as in the type. Physarum Icueophacum, 13. violascens, Rost., Mon., p. 113; Cke., Myx. Brit., p. 15. On moss. Epping Forest. (Rostafinski's Synonyms.) S2'>hacrocar2nis alhus, Bull., p. 136, var. 3, 4 (1791). Trichia filamcntosa, Trent, p. 227 (1797). Physarum eonjiuens. Link, Diss., ii., 42 (1809). Physarum connexum, Link, Diss., ii., 42 (1809). Pliysarum hy'pnorum, Link, Diss., ii., 42 (1809). Physarum alho'punctatum. Link, Herb. Physarum clavus, Ehr., Herb. Physarum conglohaticm, Ditm., t. 40 (1817). . Physarmn leucojjhaeum, Fr., Sym. Gast., p. 24 ; Syst. Myc, iii., 132 (1818); Cooke, Fung. Brit., ii., n. 519. Didymium mclanoims, fi. clavus, Wallr., non Fries (1833). Didymium tcrrestre, Fr,, in Weinm. (1836). Physarum alhipcs, De Bary, not Link (1859). Physarum striatum., Fckl., Sym. Myc, 342 (1869). Didymium licmisphcricum, Fckl., Sym. Myc, 341 (1869). Physarum granulatum, Balf. fil. (figs. 68 — 70). Sporangia stipitate, globose, sometimes slightly flattened below, grey, with small innate granules of lime, and having in 290 A Monograpli of the Myxogastres. addition a sprinkling of large, m^Jcrficial, snmv-ivhitc, amorplwus lumps of lime; stem about equalling the sporangium, from dirty grey to brown, dilated downwards, and passing into a dark hypothallus, longitudinally striate ; capillitium colourless, form- ing an irregular net, threads very thin, often flcxuous, nodes small, often flattened, only very few containing lime; spores globose, minutely warted, 9 — 10 ju diameter. Physarum granulatum, Balf., Grev., vol. x., p. 116 ; Sacc, Syll., n. 1204. (Type in Herb., Kew.) On wood. In Herb. Currey, Kew. No locality, but un- doubtedly British. About Vo mm. high, closely resembling Physarum leiicophaeum in the capillitium and spores, but distinguished by the large granules of bicarbonate of lime on the sporangial wall. In the capillitium the present species approaches the genus Tihnadoche, but differs in the great amount of lime on the sporangium wall. Physarum Phillipsii, Balf. fil. (figs. 77. 78). Sporangia stipitate or subsessile, laterally comjiressed, irregular, subreniform, subpyriform, sometimes coalescing, dark grey, generally more or less rugose, with numerous minute, white, innate particles of lime ; stem much shorter than sporangium, thick, often flattened, blackish, sometimes almost obsolete ; colu- mella absent; capillitium well developed, white, knots of lime large, irregular, numerous, conneetcd by short, thin portions; spores lilac, globose, minutely ■warted, 10 — 12 }x diameter. Physarum Phillipsii, Balf. fil., Grev., vol. x., p. 116; Sacc, Syll., n. 1208. (Type in Herb., Kew.) On manured ground in an orchard house. Shrewsbury. A very marked species, possibly having its nearest ally in Physarum nephmdcum, Rost. (I. B. B.). Mr. A. Lister has appended the following note to the type specimen of the present species in the Kew Herbarium : — " Should not this be called Physarum comprcssum, A. and S., Phymrnin. 201 as the older name? I have grown it from plasniodium in large quantity, and have sporangia answering both descriptions." The description of Physarum compressum, by Albertini and Schweinitz, is as follows : — " P. compressum. Stipitate, peridium umbilicate, compressed like a bivalve shell, obversely parabolic, grey. Stem of medium length, firm, brown. Peridium convex above, dehiscing by a longitudinal fissure. Threads of capil- litiuui rather abundant, white." Alb. and Schw., Com. Fung., p. 97. I am convinced that the present species is Phymrvm PhiUipsil, Balf. fil., the type specimen is in a good state of preservation, and the detailed description given by the author will enable any one to recognize the species. In reply to Mr. Lister's question as to whether it is not the P. compo'essicm of A. and S., I must admit that I am not at all certain; the stem is not hroivn as it should be, according to the description by the last- named authors; the white capillitium, without further ([uali- fication, is common to scores of species, therefore the only feature which remains is the compressed sporangium, and this I cannot admit as sufficient proof that Physarum Phillipsii should be reduced to a synonym of Physarum compressum, A. and S. If Mr. Lister's reasoning is sound, then we must admit that pocket-lens characters are all-sufficient for the determination of species, and that the more laborious and exact system of microscopic examination initiated by Dr. Rostafinski is a mere farce, and altogether unnecessary. In writing the above I have assumed that Dr. Lister is not in possession of the type specimen of Physarum compressum^ A. and S. Physarum didermoides, Rost. (figs. 82 — 84). Sporangia clUptic-oUong, stipitate or rarely sessile, spriiujiug from a h'oad, membranaceous, whitish hypotliallus, wall double, outer chalk-white, brittle, soon falling away; inner membran- aceous, gi'ey; capillitium with numerous, rather large nodes containing colourless granules of lime ; spores globose, brownish- violet, minutely vxtrtcd, the warts often arranged in lines ; 12 — 15 ix diameter. 292 A Monograph of the Myxogastves. Fhysaruvi didcrmoides, Rost., Mon.,p. 97, ffgs. 74, 87; Cooke, Brit. Myx., p. 11, figs. 74, 87; Sacc, Syll., n. 1171; Raunk., Myx. Dan., p. 72. Didymiiim congcdum, B. and Br., Ann. Nat. Hist. (1850), p. 865 ; Cooke, Hdbk., n. 1130. Spv.maria lichcniformis, Sz. (specimen from Schweinitz in Herb. Berk.). Exsicc. — Sacc, Myc. Ven., 269 (as Didymium conycsium, B. and Br.). On grass, twigs, bark, &c. Britain (King's Cliffe, Bulnier, Yorks. ; Carlisle) ; France ; Germany ; Italy ; Russia ; Sweden ; United States; Ceylon. Usually crowded or gregarious in considerable numbers, springing from a tough, common hypothallus ; sporangia 1*5 — 2 mm. long, stem variable in length, usually shorter than spor- angium, generally very weak, flattened or twisted. When sessile the sporangia are sometimes shorter than usual. (Rostafinski's Synonyms.) Sj)W)naria didermoides, Ach., in Pers., Syn., p. xxix (1801). Diderma oUongum, Schum., Saell., n. 1423 (1803); Flor. Dan., t. 1973, f. 1 ; Fr., S. M., iii., 103. Claustria didermoides, Fr., S. V. S., 451 (1849). Didymium congestum, B. and Br,, Ann. Nat. Hist. (1850), p. 365 ; Cooke, Hdbk., n. 1130. Didiimium farinaceum, Purt., M. F., p. 273. Physarum imitans, Racib. Sporangia hemispherical, umbilicate, greyish-white, dehiscing irregularly, stipitate, erect or slightly nodding, stem subulate, brownish-black, slightly longer than the sporangium, rigid; columella absent, capillitium white, copious, forming an irregular net, nodes irregular, sometimes filled with lime ; spores 9*5 — 10 mm. diam., violet, warted. Racib., Myx. Crac, p. 5, f, 8, a, h; Sacc, Syll, 1205. On branches. Poland, Sporangia (stem included) up to 1 nnn. hifrh. Phij.^dnun. 293 Physarum pulcherrimum, B. and R. Sporangia globose or broadly elliptical, sometimes slightly umbilicate below, stipitate, varying in colour from clear dccj) rose-red to pink; stem similarly coloured, varying length from once to twice the length of the sporangium, slender, expanding at the base into a circular hypothallus, filled with particles of lime, longitudinally wrinkled ; columella absent ; capillitium variable, sometimes scanty, at others rather copious, threads thin, forming an irregular network, colourless, or tinged pink, swollen portions elliptical, small, nuviher variaUe, interstitial, rarely produced at the nodes, containing pink granules of lime ; spores globose, reddish-lilac, very minutely verruculose, 8 — 10 [M diameter. Fhysarum pulcherrimum, B. and Rav., Grev., vol. ii., p. 65, fig. 84; Cke., Brit. Myx.. fig. 84; Sacc, Syll., vii., 1, n. 1178; Rost., Mon. Append., p. 8. (Type in Herb. Berk., Kew, n. 10,781.) On wood, &c. United States; Paraguay. A slender, delicate species, stem thin, erect; sporangium covered with a smooth crust composed of minute granules of lime. The hypothallus is usually pale ochraceous and wrinkled. Gregarious, sometimes becoming crowded, rarely 2 — 3 sporangia supjDorted on a common stem, from Vb — 2 mm. high. * Epi^porc smooth. Physarum Carlylei, Mass. (figs. 240, 241). Sporangia stipitate, broadly pyriform, orange-vermilion, mi- nutely furfuraccous ; stem about equal in length to diameter of sporangium, thick, rugulose, vermilion, expanding downwards into a small wrinkled hypothalkis ; threads of capillitium thin, yellow, forming a dense net, swollen at the angles and there containing orange-coloured granules of lime ; columella absent ; spores globose, smooth, dirty violet, 7 — 8 /x diameter. Grev., vol. xvii., March (1881)). On rotten wood. Carlisle. (Dr. Carlyle.) 294 A Monograj^h of the Muxoaastres. A very distinct species, sporangia I'o — 2 mm. high, scattered singly or in groups of two or three. Most nearly related to Physarum rnhiginosum, Fr., but readily distinguished by the smaller spores, and the scattered stipitate sporangia. Readily distinguished from the red form of Physarum ScJmmachcri, by the absence of a columella and the pyriform sporangium passing insensibly into the short stem. Physarum atrorubrum, Peck. Scattered or gregarious, stipitate ; sporangia globose, even or somewhat wrinkled, dark-red ; stem cylindrical, even, blackish or subconcolorous, capillitium when cleared of the spores whitish, sometimes with a slight pinkish tinge ; columella none ; spores globose, smooth, dark-brown in the mass, dark-red when separated, -0008— -00035' in diameter ( = about 8—9 /x). Physarum atrorulrum; Peck, 31st Report, N. York State Mus., p. 40; Sacc, Syll., vii., 1, n. 1202 (as Physarum atro- piirpurciim by mistake). Decaying wood. Adirondack Mts., U. States. The plants are scarcely one line high. The capillitium is very delicate, and when cleared of the spores the knot-like thickenings are seen to be very small and of a dark-red colour, to which probably is due the pinkish tinge sometimes observed. The dark-red granules of the sporangium walls are abundant, and appear to form a coatinuous crust. (Peck.) Physarum roseum, B. and Br. Sporangia stipitate, globose, broadly elliptical or obovate, dark reddish-jnnjile, sporangial wall covered with a thin crust of minute granules of lime; stem brownish-purple, equal, longitudinally wrinkled, filled with granules of lime, equal to or usually lunger than sporangium, expanding at the base into a small, circular, pale brown, wrinkled h}^othallus ; capil- litium well developed, threads thin, tinged pink, tvith sndtered, eUijitical, interstitial swcllinys containing lime, nodes not usually enlarged or containing granules ; spores globose, smooth, 10 — 11 m diameter. Fliys((ri(m. 295 Pliysarwm roseum, Berk, and Broome, Fungi of Ceylon, ii., n. 760, in Journ, Linn. Soc, vol. xiv,, p. 84; Rost., Mon. Append., p. 10; Sacc, Syll., vii., 1, n. 1194. (Type in Herb. Berk,, Kew.) On grass, moss, bark, &c. Ceylon ; Paraguay. Gregarious, '2o — 3 mm. high. Closely allied to Pkysarum pulchcrrimiim, of which the present species may eventually prove to be only a varfety. The spores are generally smooth under a jVth objective, but some show a minutely verruculose surface. Physarum ornatum, Peck. Sporangia depressed or hemispherical, plane or slightly con- cave beneath, greenish-cinereous, dotted with small yellow granules, the empty walls whitish ; stem short, black or Uacldsh- h'Gwn, generally longitudinally wrinkled when diy; columella none ; ca^nllitium with mimerous, yelloiv, hnot-lihe thickenings ; spores globose, smooth, violet-brown in the mass, '0004' — '0005' in diameter ( = about 10 — 11 \x). Physarum ornatum, Peck, 31st Report of N. York State Mus., p. 40; Sacc, Syll., vii., 1, n. 1220. Decaying wood. Albany, U. S. Physarum Petersii, B. and C. Gregarious, stipitate, sporangia globose, wall thin, yellow, with a thin sprinkling of minute particles of lime that soon disappears ; stem elongated, slender, subequal or slightly thicker below, expanded at the base into a small, orbicular hypothallus, rcddish-hrovm, longitudinally wrinkled and filled with lime; columella subglobose or shortly cylindrical, whitish ; capillitium rather dense, forming an irregular network, threads thin, with scattered, elliptical, interstitial swellings containing yellow granules of lime, nodes rarely swollen ; spores dirty lilac, smooth, 7 — 8 /m diameter. Physarum Petersii, B. and C, Grev., vol. ii., p. GO; Rost., Mon., p. IIG, Append., p. 6 ; Sacc, Syll., vii., 1, 1174. 29G A Monograph of the Myxogastres. (Type in Herb. Berk.) Physarurii galhcum, Wingate, in E. aud E., N. Amer. Fung., 2491. On moss, wood, &c. U. States. Resembling in the long, slender stem, and small, elliptical lumps of lime, some species of Tilmadoche from which genus it differs in the presence of a columella. About 2 mm. high. Physarum hians, Mass. Sporangia stipitate, yellowish-red, then blackish, globose or slightly elongated ; stem reddish-brown, sulcate, expanding at the base into a small, circular hypothallus, penetrating the sporangium as a columella which reaches nearly to the apex ; threads of capillitium yellowish, with violet tinge, forming a somewhat dense network, nodes irregularly oblong, reddish- yellow, spores violet, smooth, 8 — 10 /x diameter. I'ilmadoche hians, Rost., Mon. App., p. 14. Tilmadoche minuta, Berl,, Sacc, Syll., vii., 1, n. 1251. On dead leaves. Belgium. The present species is removed from Tilmadoche on account of the columella. Physarniii rufibasis, B. and Br., quoted by Rostafinski as a synonym under the present species is quite distinct, hence the Ceylon habitat of the latter is incorrect. (Rostafinski's Synonyms.) Craierium mimitum, Kickx, non Fr. Physarum ritjihasis, B. and Br., Fung, Ceylon, n. 762 (1873). Chondnoderma infiatum, Rost., Mon., p. 425, n. 177 (1874), See Kickx's description of Craterium minutum under Cra- terium. Physarum Leveillei, Rost. Sporangia lenticular, fuifuraceous, golden, as is also the upwards attenuated stem, threads of capillitium vanishing ; spores globose, smooth, 10 — 11 ij. diameter. Physarum. 297 Physarum Zeveillei, Rost., Mon. App., p. 7; Sacc, Syll., vii., 1,11.1177. On dead leaves. Chili. Var. chryso2yiijS, Rost., stern and qiorangiuiii goldcn-ycUow, 1 mm. diameter; spores 10 /x diameter, lime granules golden- yellow. Var. auri2)cs, Rest. ; stem golden, sporangium eitrin. Venezuela. Var. Fuistingii, stem orange, sporangium rii/ous-hrown. Westphalia. (Rostafiuski's Synonym.) Physarmn cmreurn, ^. ehrysopns, Lev., Ann. Sc. Nat., p. 116 (1846). Physarum Kalchbrenneri, Mass. (n. sp.). Scattered, stipitate, sporangia spherico-depressed, umbilicate below, pale yclloiu-green, rough with minute particles of lime ; stem elongated, thickest heloiv or fusiform, pale 2^'inJx;, rigid, filled with granules of lime; capillitium forming a very dense, irregular network, nodes numerous, large, irregularly angular, containing yellow granules of lime, internodes 6 — 9 ju. thick, colourless, without lime, collapsing ; spores dingy-lilac, smooth, 11 — 13 /x diameter. (Type in Herb., Kew, along with Tilmadochc varialilis.) On wood. Cape of Good Hope. Superficially resembling a typical Tihnadoehe in the long, slender stem, but separated from that genus by the very thick capillitium tubes, and the numerous large, irregularly angular nodes. From 2-5 — 3 mm. high. Physarum globuliferum, Rost. Sporangia globose, stipitate, greyish ; stem equal to spor- angium or twice as long, rigiel, fragile, lohite, suleede ; eolumeUa large, cyliiulrical, ohtuse, white ; capillitium well developed, rigid, not collapsing, vesicles containing lime numerous, of variable size, yellowish or reddish; spores globose, smooth, 9 — 11 /oi diameter. 298 A Mo7io(jraph of the Myxogastres. Physarmn glolndifcritia, Rost., Mon., p. 98 ; Sacc, Syll, vii., 1, n. 1173. On moss, leaves, rotten wood, &c. Sweden ; France ; Ger- many. (Rostafinski's Synonyms.) Sj)haerocarpus glolndifcr, Bull., Champ., j). 134, t, 484, f. 3 (1791). Stemonitis globulifcra, Gmel., Syst. Nat., ii., p. 1467, n. 2 (1791). Phymrurii glohvMfcrum, Pers,, Syn., p. 175, t. 3, f. 11, 12 (1801). Trichia globulifera, D.C., Fl. Fr., 2, p. 253 (1805). Diderma globuliferum, Fr., I.e., iii.^ p. 100 (1829). B. Sporangia sessile, j^^'^sinGdiocarj), o?' aethalioid. * Epis'iwre warted. Physarum cinereum, Rost. (figs. 71 — 73). Sporangia sessile on a broad base, sometimes globose, scattered or crowded, at others elongated, flexuous and depressed, wall rigid with lime, white ; columella absent ; capillitium abundant with numerous la7'ge, irregular knots of lime connected by very thin threads; spores globose, dingy violet, niimttely warted, 10 — 13 fj. diameter. Physarum cinereum, Rost., Mon., p. 102, figs. 71, 72, 85 ; Cooke, Myx. Brit., p. 13, figs. 71, 72, 85; Sacc, Syll, n. 1189; Schroeter, p. 128; Raunk., Myx. Dan., p. 75, t. 4, f. 9. Exsicc. — Ellis and Everhart, N. Amer. Fung., ser. II,, n. 2085 ; Rav., Fung. Car., 79 and 83. Fung. Cub. Wrightiani, 535; De Thum., Myc. Univ., 1598; Holl, Schmidt and Kunze, Deutsch. Schw., 38 ; Roum., Fung. Gall., 1881. On leaves, wood, twigs, &c. Britain (King's Cliffe, Bury St. Edmund's, Bungay, Kew, Hitchin, Carlisle, Scarboro', Berwick, N. Wales, Linlithgow) ; Germany; Fhdand; Sweden; Belgiimi; PJnjmrum. 299 France; Switzerland; Algiers; S. Africa; United States; Ceylon ; Australia ; India. Varying from -5—2-3 mm. when elongated ; some forms of the present species resemble externally Didyminm scrpula, Fr., but are readily distinguished under the microscope. Var. ovoidcutn, Sacc. Sporangia vertically ovoid, densely gregarious. riiymruvi cinercum, var. ovoidcuvi, Sacc, Mich., ii., p. 334. Britain (Lyndhurst, New Forest) ; France. (Rostafinski's Synonyms.) ^ Mncilago crusfacca, Mich., Gen., t. 96, f. (1729). Lycoperdon cincreum, Batsch, fig. 1G9 (1783). Lycoperdon alni, Bjer., in Vet. Handl, p. 39 (1789). Trichia caerulea, Trent., p. 229 (1797). Physarum cincreum, Pers., Syn., p. 170 (1801) ; Nees, f. 107; Let ell., t. 710, f. 2. Fhysaruvi violaccnm, Schum., Saell., 1428 (1803) ; Fl. Dan., t. 1980, f. 2. Fhysarum corrugatum, Link, Herb. Physartt77i coelahim, Ehr., Herb. Fhysarum conglolatum, Fr. Gast., p. 21 (1818); Letell., t. 710, f. 3. Didymium cincreum, Fr., S. M., iii., 126 (1820); Eng. Fl., v., p. 314; Cke., Hdbk., 1133. Fhysariim plumhcum, Fr., S. M., iii., 142 (1829). Fhysarum Weinmanni, Fr., var. in litt. Didymium melanoims, Wallr., n. 2193 (1833). Fhysarum sinuosum, Wallr. Didymium scrohiculcdum, Berk., Hook. Journ. (1845), p. ijQ>. Fhysartim album, Fuckel, F. Rhen., 1469 (1865). Var. ficxuosum, sessile on a broad base, depressed, spherical in outline, or elongated and variously curved, wall at first white, becoming grey when the outer chalky layer has broken away in flakes; columella ab-scnt; capillitium dense, threads thin, 2 ju diameter, culnirless, Jlcxuous or spindly coiled, furnished 300 A ]\/ono{/raj)h of the Mifxiyjastres. witli nuniermos small, discoid, coloured 2'>'>'oj<-'<'tions irregularly scattered, combined to form an irregular network, intcrstiticd, small, ellipticcd siocllings containing colourless gramdes of lime, very rare ; spores brown witli lilac tinge, globose, coarsely tcartcd, 14 /u, diameter. Physariim cinerettm, Sacc, Myc. Ven., Exs., n. 1101. On living or dead leaves, twigs, &c. Britain (Epping Forest) ; Italy. The above, if in reality a variety of Pliysarwn cinereum, differs considerably in the structure of the capillitiiini and the large, coarsely warted spores. Its occurrence in Britain and Italy suggests it to be a constant form. Physarum chrysotrichum, B. and C. Sporangia sessile, subglobose when scattered, often irregular or crowded and more or less confluent, wall rigid, usually granulated with 'projecting lum'ps of lime, varying from yellow through fulvous to almost clear vermilion; capillitium dense, nodes mtmcrous, large, irregularly angidar, filled loith yellow or taiuny lumjps of lime, connected at various jwints by long, thin, cm2Jty internodes, some of the nodes are small and without lime ; spores lilac, globose, mimttely warted, 7 — 8 /x diameter. Physarum chrysotrichum, B. and C, Grev., vol. ii., p. QQ. (Type in Herb. Berk., Kew, n. 10,804.) Physarum inacqualis. Peck, 31st Report, p. 40. Badhamia decijnens, Host., Mon. App., p. 4. "Physarum suhglohosum, (B. and C.)." Sacc, Syll., vii., 1, n. 1226. (I am not aware that there is any such species by B. and C, and it is obviously a slip in Saccardo's Sylloge for P. chrysotrichtiin. On moss, wood, &c. United States. The type specimen is a true Physarum, as will be seen by the above description, and on the same piece of moss-covered wood ranges through the colours given above. Physarum scrobiculatum. Mass, Sporangia sessile on a broad or narrowed base, seated on a Phiisarnm. 301 thick, vAde-sjrreadinff Jn/jwthallns, scattered or aggregated, often irregular and aetlialioid, or several blending together to form a plasniodiocarp, ^:>a/(; JlcHli-colour, brittle, often furfuraceous ; capillitium dense, forming an irregular network, nodes numerous, large, irregularly angular, filled with yellow granules of lime, connected at several points by long, thin, colourless internodes without lime ; columella absent ; spores lilac-brown, globose, minutely loarted, 13 — 14 /x diameter. Didyminm scrobiculatum, Berk., Hook. Journ., vol. iv., p. 60. Physariim cinerenm, Sacc, Syll., vii., 1, n. 1189 (in part.). (Type in Herb. Berk., n. 10,774.) On charred wood. Australia. Distinct from P. cincreuvi in the size of the, spores, wide- spreading, firm hypothallus, and colour of the sporangial wall, Physarum Rostafinskii, Mass. Sporangia distinct, conglomerated, sessile, rounded or angular, depressed, exterior membrane thickish, granular, chalky, fragile, yellow or ochraceous, interior wall thin, grey or yellowish, from 0'2 — 0"5 mm. diameter; capillitium furnished with numerous irregularly angular nodes containing coloured granules of lime ; columella cylindrical, spores minutely spinulose, blackish-violet, 8 — 9 jm diameter. Physarurii eonglomeratum, Eost., Mon., p. 109; Sacc, Syll., vii., 1, n. 1184. On leaves, moss, &c. Germany; Sweden; Finland; San Francisco. I am not acquainted with the present species, which is certainly not Biderma eonglomeratum, Fr., as supposed by Rostafinski, hence the localities given by Rostafinski may or may not show the distribution of this species ; the same applies to his synonyms. (Rostafinski's Synonyms.) Biderma ochraceum, Hoffm., Fl. Germ., iii., t. 9, f 2, h (1795). Ecticularla ochracca, Poir., Ency. ap Streintz (1795). 302 A Moiio. thick, combined to form a loose, irregular network, nodes very rarely slightly in- crassated, and containing a few minute, colourless granules of lime; spores globose, dirty lilac, smooth, 10 fj. diameter. On wood. Venezuela. An apparently very distinct species, remarkable for the almost total absence of thickened nodes or swollen interstitial portions of the capillitium tubes, hence approaching the section Hemi- didymium of Didymium. From 3 — 3"5 mm. high. Tilmadoche oblonga, Kost. Scattered or gregarious, stipitate ; sporangia cylindrical, obtuse at both ends, apex hroadly and deeply umhilicate, tawny-olive, sprinkled with pale particles of lime ; stem elongated, slender, becoming thinner upwards, straight, or usually more or less curved near the apex, reddish or yellow-brown, more or less longitudinally wrinkled, filled with minute particles of lime ; capillitium well developed, threads colourless, thin, with scat- tered, small, interstitial, fusiform swellings containing colourless or yellowish granules of lime, angles of bifurcation not in- crassated ; spores dingy lilac, globose, mimitely vcrrucidose, 7 — 9 [i diameter. Tilmadochc ollonga, Rost., Mon. App., p. 13; Sacc, Syll., vii., 1, n. 1250. Trichamphora oblonga. Berk, and Curtis, Grev., vol. ii., p. QQ ; N. Amer. Fung., n. 360. (Type specimen in Herb. Berk., Kew, n. 10,917.) Physarella mirahilis, Peck, Bull., Torr. Bot. Club., vol. ix., p. 61, pi. 24, figs. 1—6 (wretched figures); Rev. Myc, 1882, p. 172, tab. xxix., fig. 5 (after Peck's figures) ; Sacc, Syll., vii., 1, n. 1227. Exsicc.—EW'is, N. Amer. Fung., n. 1212. On wood, leaves, &c. United States, Tilmadoclie. ^ 335 About 2 mm, high, stem straight or curved, thin, sporangium typically cylindric-oblong, broadly and deeply umbilicate at the apex; there is no trace of a columella. The present species appears to be inclined to sport. Mr. Harold Wingate, of Philadelphia, has sent me a fine series of forms of the present species passing from the typical form described above to speci- mens that are much larger, with a shorter, thicker, red stem, which is in some instances branched and bearing 3 — 5 sporano^ia. These vary in form from the typical condition, through cyathi- form to broadly funnel-shaped, with a recurved, often flexuous margin, externally reddish-brown, internally orange-yellow; in these forms the umbilicus has become so deeply depressed that the cavity of the sporangium is almost obliterated. It is the inner depressed apical portion of the sporangium that Peck describes as a spurious, hollow columella; he has obviously formed the genus Physardla from an abnormal condition of the present species. Tilmadoche gyrocephala, Rost. Sporangia irvcgnlarly lobcd or lacimosc, often compressed and umbilicate below, stipitate, wall thin, covered with irregular yelloio or greenish-yellow scales of lime ; stem elongated, tapering upwards, weak and often curved, irregularly rugulose, passing downwards into a thin, wrinkled hypothallus, bright yellow or orange ; columella absent ; capillitium well developed, formino- a loose net, nodes elongated, filled with yellow granules of lime, internodes long, thin; spores globose, minutely wartcd, dingy, lilac, 7 — 9 ii diameter. Didymimn gyroce-phalwm, Mont., Ann. Sci. Nat., Ser. II. vol. viii., p. 362; Mont., Syll, n. 1073. Tilmadoche gyrocc^Jhala (Mont.), Rost., Mon., p. 131; Sacc, Syll, vol. vii., pt. I., n. 1248; Mass., Journ. Myc, vol. v., p is/ t. 14, f. 8 (1889). Oribraria staminiformis, Speg., Fung. Arg., Puo-. II., n. 109. ^xsicc.—Rav., Fung. Amer., n. 477 (as Physarum ScJinmachcri, Spr.). 336 A Monograph of the Mi/xogastres. On twigs, moss, &c. Brazil ; Argentine Republic ; Aiken, S. Carolina. Gregarious, 1"5 — 2 mm. high, having the habit of rinjsarum Schumacheri, but readily distinguished by the absence of a columella and the very irregular sporangia which is not due to the coalescence of several individuals, as proved by the thin, simple stem. Tilmadoche reniformis, Mass. (n. sp.). Gregarious ; sporangia reniform, deeply umbilicate below, some- times umbilicate above also, and then presenting a dumb-bell- like form, coating of lime white, often becoming cracked, stem elongated, slender, erect, straight, tapering upwards, pale brown, wrinkled, filled with granules of lime ; capillitium rather scanty, threads forming a very loose net, with scattered, fusiform swell- ings, colourless or tinged yellow ; spores globose, dull purple, coarsely spinulose, 16 — 17 /x diameter. Ceylon. (Type in Herb. Berk., Kew.) On bark. Gregarious, rarely scattered, 2 mm. high. The sporangium is sausage-shaped and curved, the stem entering the concave side or umbilicus. This peculiarity, and more especially the large, coarsely spinulose spores, characterize the species. Ceinkowskia, Rost. Wall of sporangium single, containing lime, dehiscing irregu- larly; capillitium attached at many points to wall of spor- angium, forming an irregular network having numerous free, curved, pointed ends. CeinkowsJcia, Rost., Mon., p. 91; Cooke, Brit. Myx., p. 11 ; Schroeter, p. 181 ; Sacc, Syll., vol. vii., pt. I., p. 329. Physarum, Alb. et Schwz., p. 90. Didcrma, Fr., S. M., iii., p. 112. Didymium, Bischoff. Kryptogamenkunde, f. 361. Ceink<)iv>mi Ulaciiwm, x 50. „ .308. Spore of same x 1200. „ 309. Oraterinm rnhiginosiim, x 50. „ 310. Spore of same, x 1200. ,, 311. Retimlaria lycoperdon, a. castaiieu.ni ; nat. size. „ 312. Spore of same, x 1200. „ 313. Clwndriodcrma Michelii, a, hypothallus, x 50. GENERA AND SPECIES EXCLUDED. Endocalyx, B. and Br., belongs to the Fungi. Reticularia affinis, B. and C, -\ „ atro-rufa, B. and C, - = Trichospormm Curtisii, „ venulosa, B. and C, i ^''''■' ^^'''''- ^^^^^ ^^^^- ^ . . , , ^ 1 r Trichosporium phyrrho- Reticiilariapliyrrliospora,Berk.J ^^^^.^^^ ^^^^^^ j^^^.^ „ rubra, Ayres. ( ^^^^^ ^gg^ {Trichosporium apio- = sporium, Mass., Journ. Myc, 1889. Badhamia fulvescens, Cke,, is probably a member of the Ferisjjoriaceae. Didymium bulbillosum, Br. and Br., = minute galls on a leaf, as proved by examination of type. BIBLIOGRAPHY. Baranetzki ; Influence de la lumiere sur Ics plasmodia des Myxomycetes. Mem. Soc. Nat. de Sci. Nat. de Cherbourg, v., xix, Bary, De ; Morphology und Physiologic der Pilze, Mycetozoen und Bacterien; Leipzig, 1866 (English edition, Clarendon Press, 1887). ; Die Mycetozoen ; Zeitschr. f. wissensch. Zool, Bd. x., 1859. Bibliv(jv(ip}nj. 357 Brefeld; Didyostelmm'mucoroidt's ; Abhandl. Jer Senk. Nat. Gesell., vii., 1809. Brunchorst, T. ; Sjion.fpsjwra Solani (a new Myxomycete causing disease in potato) ; Bergens Museums Aarsbcret, 1877 (2 PI.). BuUiard ; Hist, des Champignons. Candolle, Le ; Flore Frani^aiso ; fungi in vol. ii. Cienkowski, L. ; Entw ick. der Myxomycetcn ; Pringisli. Jalirb., iii. ; Giittulina rosea, Bot. Jahresb., 1873, p. 61. ; Zur Entwickel. der Myxomyceten; Jahrb. fur Wiss., Bot. iii., 1862. ; Das Plasmodium; Jahrb. fur Wiss., Bot. iii., 1863. Cooke, M. C. ; Myxomycetes of Great Britain, 1877. Crouan, P. L., and H. M. ; Florule de Finisterre. 1867. Dangeard, M. P. A. ; Recherches sur les organisms inferieurs ; Ann. Sci. Nat. (Bot.) iv. (1886), (4 PL). Fayod, V. ; Beitrag zur Kenntniss niederer Myxomyceten ; Bot. Ztg., 1883. Fries, E. ; Systema Mycologicum ; vol. iii. (1829). '; Stirpium agri Femsjonensis (1825, 1826). ; Summa Vegetabilium Scandinaviae (1846). Fuckel, L. ; Symbolae Mycologicae (1875, concluded). Gobi, C. ; Fscndospora, structure and development of; Ber. Gesell. off Gesundh.; Petersburg, 1887 (in Russian); (abstract in Bot. Centralbl., 39 (1889), p. 346. Greville, R. K. ; Flora Edinensis (1824). ; Scottish Cryptogamic Flora, 1823—1829. Henfrey; Notes on the elaters of Trichia ; Trans. Linn. Soc, v., 21, p. 221. Hisinger, E. ; Tctramyxa imo'asitica (producing tubercles on mxiina rostellata and zanichcllia jjolycarpa). Meddl. Soc. pro Faun, et Flor. Fennica, 1887. Lister, A ; Notes on the plasmodium of Badhamia utrknlosa and Brefeldia maxima; Annals of Botany, vol. ii., 1888 (2 PI.). ; Notes on Chondriodcrma difformc and other Mycetozoa ; Ann. Bot., vol. iv. (18911) (1 PI.)- 358 A Monograph of the Myxogastres. Lister, A. ; New species of Mycetozoa ; Joimi. Bot., 1801 (4 PI.). Massee, G. ; A revision of the Trichiaceae ; Journ. Roy. Micr. Soc, 1880 (4 PI). ; Mycological Notes; Journ. Mycol., vol. v., 1880. Persoon, D. C. H. ; Observationes Mycologicae, 170G — 1700. ; Synopsis Methodica Fungorum, 1801. Raciborski, M. ; Myxomyceteu der Tatra; Hedw., 1885, p. 168. ; Bemerk. liber einige in den letzten Jaliicn besehrieb. Myxomyceten ; Hedw., 1887, p. 100. Raunkier, C. ; Myxomycetes Dauiae ; Bot. Tidsskr. (1888 — 1880). Rex, G. A. ; A remarkable variation of Stemonitis Baucrlinii, Massee ; Proc. Acad. Nat. Sci. Phil., 1800, p. 36. Rosanoff; De I'influence de I'attraction terrestre sur la direc- tion des Plasmodia des Myxomycetes ; Mem. Soc. Imp. Sc. Nat. Cherb., vol. xiv. Rostafinski, J. ; Versuch eines Systems der Mycetozoen (Inaugural Dissertation), Strassburg, 1873. ; Sluzowce (Mycetozoa), Monogratia (In Polish, numer- ous figures) ; Paris, 1875. — Supplement to above, Paris, 1876. Roze, E. ; Des Myxomycetes et de leurs place dans la systeme. Bull. Soc. Bot. France, vol. xx., p. 320 (1873). Saccardo ; Myxomycetes ; in " Sylloge Fungorum," vol. vii., pt. I. Schroeter, J. ; Myxomycetes, in Cohn's Kryptogamen — Flora Von Schlesien, 1885. Schumacher, C. F. ; Enumeratio Plantarum Saellandiae, 1801. Schweinitz, L. D. de ; Synopsis Fungorum- Carolinae; Soc. Nat. Curio. Lips., 1882. ; Synopsis Fungorum in America Boreali media deg- cntium. Trans. Amer. Phil. Soc, vol. iv., New Ser. (1834). Stahl, E. ; Zur Biologie der Myxomyceten ; Bot. Ztg. (1884), 145, 146, 161—176, 187—101. Tieghem, Ph. Van; Traite de Botanique, 1884. ; Sur queLjucs Myxomycetes a plasmode agn'ge ; Bull. Soc. Bot., xxvii., 1880. HihUoin-apliii. 359 Wallroth, F. G. ; Flora Cryptogamica Germaniae, 1833. Ward, H. M. ; Tlio Morphology and Physiology of an aquatic Myxomycete; Studies from the Biological Laboratories of the Owens College, vol. i. (1886). Wigand; Zur Morphologie der Gattungen Trichia imd Ar- cyria; Jahrb. fur. Wiss. Bot., iii. (1862), Wingate, H. ; Orcadella, a new genus of Myxomycetes ; Rev. Mycol. (1800), p. 7-1, 75. Woronin and Famintzin ; Ceratium Hydnoidcs and C. •pwoidcs. Mem. de I'acad. Imp. des. Sc. de St. Petersb., vii., ser. T., n. 3, 1873. Woronin; Plasmodiophora Brassicae ; Jahrb. fur. Wiss. Bot.. xi., p. 548 (1878). ZofF, W. ; Vorkommen von Fettfarbstoffen bei Pilzthieren. (Mycomyceten) ; Flora; Ixxii. (1880), pp. 353—361 (1 fig.). ; Ueber Pilzfarbstoffe (including some of the Myxomy- cetes) ; Bot. Ztg., xlvii. (1880), 54—61,^00—81, 85—02 (1 PI). ; Die Pilzthiere oder Schlcimpilze ; in Schenk's " Hand- bucli der Botanik," parts 15, 16 (1884), Zukal, H, ; Einig neue Pilze, Myx. und Bact. (contains de- scriptions of two new Myxogastres, Trichia nana and Arnau- rochadc spcciosa) ; Verhandl. K. K. Zool. — Bot. Gesell. Wien., XXXV. (1885), pp. 332—345 (1 PI). INDEX OF GENERA. Bynomjms arc printed in Italics. Alv-Lsia, B. and Br., 126. Amaurochaete, Rost., 89. Ancyrophorus, Raiuik., 107. Arcyria, Hill., 141. Badhamia, Berk., 316. Brefeldia, Rost., 90. Chondrioderma, Rost., 197. Chromdsjinrhnii, Sacc, 93. Cienkowskia, Rost., 336. Clathroptychium, Rost., 50. Omiidrirhd, Preuss., 72. Conntria, Rost., 126, 130. Crateriachea, Rost., 344. Craterium, Toent., 262. Cribraria, Pers., 54. Dcnnodlnm, Rost., 120. Diachaea, Fries., 258. Dictydium, Sclirad., 67. Didymium, Schrad., 218. Echinostelium, De Bary, 108. Enerthenema, Bowin., 104. Enteridium, Rost., 44. Fuligo, Rost., 339. Hernia rcyria, Rost., 141. Hetcrodictiion, Rost., 54. Heterotrichia, Mass., 139. Lachnobolus, Fries., 136. Lamproderma, Rost., 94. Iieocarpus, Rost., 337. Lepidodex-ma, De Bary, 251. Licea, Schrad., 35. Lindbladia, Fries, 35. Lycogala, Midi., 119. Lycojjcrdon, Bux., 120. Oligonema, Rost., 169. Ophiotheca, Curr., 130. Orcadella, Wiii!^., 48. Orthotrichia, Wing., 109. Perichaena, Fries., 113, 130. Physaruni, Pers., 273. Frotuderma, Rost., 43. Protodermium, Ro.st., 43. Prototrichia, Rost., 126. Raciborskia, Berl, 108. Reticvilaria, Bull., 92. Rostafinskia, Speg., 91, 108. Siplioptycbium, Rost., 88. Spumaria, Pers., 256, Stemonitis, Gled., 72. Tilmadoche, Rost., 327. Trichia, Haller., 126, 173. Tubulina, Pers., 35. INDEX OF SPECIES. ASynonyms arc in roinan ty]_)C. Ahrupta (Trich.) 187. acmnliKifa (Stem.) 78. Adnata (Arcy.) 145. advennla (Trich.) 181. aeneuin (Olig.) 172. o^ne (Phys.) 283. affi,)ie (Chon.) 210. njjiue (Didy.) 222. affinis (Coma.) 77. oijinh (Arcy.) 157. dfiinis (Trich.) 194. affinis (Stem.) 77. alha (Spurn.) 256. albescens (Chon.) 209. albescens (Dider.) 209. Albicans (Fhjs.) 312.^ Alexandrowiczii (Chon.) 212. AlexcuKlroroic'di (Badh.) 324. AlexandrovAczii (Didy.) 232. angnlatmn (Didy.) 249. anomala (Tilm.) 333. anomala (Ophi.) 135. anomala (Trich.) 135. anomala (Corn.) 135. anumalnm (Didy.) 245. anomalum (Chon.) 245. antiades (Trich.) 314. antiades (Sphaer.) 314. antiades (Pliys.) 313. applanata (Retic.) 45. applanata (Peri.) 116. applanata (Hemi.) 117. arojrclla (Lach.) 138. arcyrioides (Lamp.) 95. arcyrioides (Lamp.) 102. arcyrioncma (Lamp.) 96. aiijillacca (Crib.) 65. argillacea (Crib.) 37. artocreas (Peri.) 135. atra (Stem.) 78. cdra (Amaur.) 89. atropurpureum (Lyco.) 121. atroruhrum (Phys.) 294. atrum (Phys.) 314, 315. aurantiaca (Arcy.) 155. avrantiaca (Crib.) 57. anreum (Crat.) 269. australe (Steg.) 119. australis (Rost.) 92. austrulis (Didy.) 237, aiistralis (Peri.) 119. Balfouri (Trich.) 186. Barteri (Didy.) 231. Bauerlinii (Stem.) 79. Bavarica (Olig.) 170. Bavarica (a) 170. Berkeleyana (Ener.) 106. Berkeleycumm (Clion.) 214. Berkeley i (Phvs.) 285. Berkeleyi (Pliys.) 242. Berkeley i (Clat.) 53. Berkeleyi (Tilm.) 332. bicolor (Arcy.) 153. bieniasdi (Crib.) 60. bieniaszii (Heter.) 61. bornharda (Proto.) 128. Bonarioisis (Arcy.) 157. botryoides (Didy.) 229. botrytis (Trich.) 176. Braunianum. (Phys.) 30S. brevijilmn (Olig.) 173. Broomef (Olig.) 172, brunuea (Tubu.) 42. brunnea (Licea) 42. brunncolum (Phys.) 280. 362 A Monograph of the Mi/jeogastres. brunneolum (Dider.) 281. BucknaUi (Arey.) 161. Buckmilli (Heuii.) 161. caespitosa (Tiibu.) 43. cafspitosum (Pliys.) 43, 311. cdlcarenm (Clion.) 218. calyndata (Arcy.) 162. calyculata (Hemi.) 162. candidum (Pliys.j 286. cnnojiavescens (Ophi.) 134. Capoise (Phys.) \309. capsulifera (Badli.) 319. Cdfcstidnuin (Lepi.) 255. Cdrr.stiaiia (Rcti.) 255. Varhjlmna (Tricli.) 174. Carlylei (Pliys.) 293. Carlylei (Stem.) 84. CarmdchaeUanum (Chon.) 202. Carmicliaelianum (Dider.) 202. Caspnvfii (Siph.) 89. C'(viprs\Ti\m.) 333. cervhi-iinim (Fliys.) 306. ccnatum (Diet.) 67. ChailetUi (Lepi.) 255, Chamaeleontina (Proto.) 130. chondroderma (Didy.) 232. chrysopeplum (Phys.) 275. clirysosperma (Ophi.) 131. chrysosperma (Trich.) 189. chrysospora (Arcy.) 164. chrysospora (Ilenii.) 164. Chrxjsotricimm (Phys.) 300. cinerea (Arcy.) 151. cinerea (Arcy.) cribroides (Var.) 153. cinereum (Phys.) 301. cinereum (Phys.) 310. cinereum (Phys.) 298. cinnaharina (Licea) 52. cinnahariu.um, (Chit.) 53. clnnammnea (Arcy.) 158. circinans (Lach.) 139. ciicumscissa (Corn.) 131. rirriim.scissa (Ophi.) 131. cit rlnelh wi (Phys.) 278. citriiium (Phys.) 275. citrimun (Phys.) 278. davafa (Arcy.) 165. clavata (Hemi.) 166. damts (Didy.) 230. coaihiata (Badli.) 325. cohnnhhin (Tilm.) 331. cohiuil)innm (Phys.) 332. culumbinum (Didy). 331. confliiens (Coma.) 77. conflnens (Didy.) 235. cotijliisiis (Stem.) 77. canfnsa (Peri.) 117. covfnsa (Diac.) 259. fonfnsnm (Crat.) 263. colicmbhmin (Lamp.) 100. commutable (Didy.) 229. compada (Tihn). 332. complanatum (Didy.) 234. compresseum (Phj^s.) 286. concinnum (Dider.) 308. coiiciiiniim (Phys.) 308. congesta (Lach.) 139. congestiim (Didy.) 292. conglomeratum (Phys.) 301. conglomeratum (Phys.) 304. conglomeratum (Dider.) 301. conicum (Dermo.) 123. conicum (Lyco.) 123. connatum (Didy.) 251. contextiim (Phys.) 303. contoHa (Trich.) 182. Cookei (Didy.) 245. Cookei (Chon.) 245. Cookei (Chon.) 207. Cookei (Arcy.) 154. corticaUs (Peri.) 115. corfcicalis (Peri.) 135. crassipes (Ancy.) 107. cribrarioides (Stem.) 104. cribroides (Var.) 144. croceo-flavium (Didy.) 277. crndacenm (Chon.) 215. crustaceum (Dider.) 215. crustaeeum (Didy.) 235. ernstacevm (Dider.) 215. cubense (Chon.) 213. cuprca (Proto.) 129. cup)ripeii (Phys.) 284. Curtisii (Didy.) 272. Gnrtmi (Crat.) 272. Curtisii (Badli.) 272. Curtisii (Scyp.) 272. cylindrica (Tubu.) 39. cylindrica (Licea) 53. cylindricum (Crat.) 268. daedaleum (Didy.) 250. dealbata (Chon.) 207. dealbatum (Didy.) 208. Dccaisnemia (Trich.) 185, decipiens (Arcy.) 158. decipiens (Arcy.) 157. Index. 3G3 decipions (Badli.) 300. depliiiiJituin (Clion.) 215. drpirssii (Peri.) 114. dldiidloidrs (Crib.) 65. dictyociirpu (Corn.) 131. didi/onoiia (Arcy.) 154. dktuuspennvm (Crut.) 270. dictyospora (Badli.) 271. dictyospora (Stem.) 83. diderma (Phys.) 304, didcrmoidrs (Pliys.) 291. diffonne (Clion.) 213. dipitata (Stem.) 153. dUjitata (Arcy.) 153. discoideum (Didy.) 229. dissllien.s (Clat.) 53, dissiliens (Diet.) 54. Ditmari (Pliys.) 277. dubiiim (Didy.) 244. dubiiim (Didy.) 246. eehinoHpora (Didy.) 239. echiniilata (Stem.) 97. echinidatiim (Lamp.) 97. effusa (Lind.) 41. efum (Tubu.) 41. cfvsum (Pliys.) 313. ctf'iisntii (Didy.) 236. data (Crib.) 61. clecjam (Pliys.) 313. elecians (Crib.) 55. elegants (Euer.) 105. elegans (Raci.) 108. elegantissimum (Didy.) 243. ele-antula (Proto.) 128. eUipsospoiiim (Pliys.) 310. milsi(n,<( (Lamp.) 98. cpidendrum (Lyco.) 121. ereda (Trich.) 184. erytliriniun (Didy.) 275. arythrinum (Didy.) 249. exigium (Clion.) 240. eximium (Didy.) 241. Fainnani (Didy. ) 249. I'allax (Licea) 47. faUax (Chon.) 212. fallax (Trich.) 192. Famintzini (Phys.) 309. farinacenm (Didy.) 219. 'farinaceiim (Dider.) 233. fascicidafa (Padli.) 326. iasciculata (/I) 163. iascieulatum (f.) 230. fasciculatum (f.) 288. ferrvginea (Arcy.) 144. ferriiiihwa (Stem.) 85. Jimdariinii (Pliys.) 314, Jhuidlifer (Proto.) 127. ilagellifer (Derm.) 127. lia-elliter(Tricli.) 127. flagellifera (Proto.) 128. flavicommn (Didy.) 242. fiavicomum (Phys.) 242. liavicomnm (Phys.) 285. flavida (Peri.) 171. flavidum (Didv.) 251, flavidiim (Didy.) 251. flavidum (Phy.s.) 247. flavidum (Didy.) 247. flavidum (Olig.) 171. flavo-fnscum (Lyco.) 124. flavu-virens (Phys.) 310. flaviim (Phys.) 279. flexuosa (Tubu.) 37. flexuosa (Licea) 37. floriforme (Chon.) 308, 198. fluminoitiis (Stem.) 80. 'fragilis (Spha.) 176. frag His (Trich.) 175. fragilis (Trich.) 176. Friesiana (Coma.) 82. Friesiaiia (Stem.) 82. Friesiana (Peri.) 135. Friesianum (Chon.) 208. Fricsii (Crat.) 266. Friesii (Arcy.) 152. Fvckdiana (Badh.) 321. Fuckeliana (Trich.) 321. Fuckdiannm (Didy.) 222, Fiidielia)nim (Lamp.) 103. Fnckelii (Crat.) 272. fuliginea (Hemi.) 169. fidiginea (Arcy.) 169. fulvdla (Badh.) 326. fulvellum. (Didy.) 237. fidvipcs (Didy.) 248, fidcmii (Lepi.) 253. fumdiiw. (Olig.) 173. fnsca (Stem.) 72. I'usca (Stem.) 88. fusca (Arcy.) 149. fnsco-atra (Peri.) 115, GahrieUae (Hetev.) 140. galbcnm (Phys.) 296. gcastroides (Chon.) 201. geastroides (Dider.) 201. 364 A Monograph of the Myxogastres. glaucnm (Didy.) 284. (jlavcnm (Phys.) 284. '(llobosmn (Chon.) 206. qlohofoi.^ (Lach.) 137. illohidlfennn (Pliys.) 297. yracih'vta (Tiliii.) 330. (jmnulatvm (Phys.) 289. fjranuUfcra (Badh.) 321. graniiliferum (Didy.) 321. Guaranitica (Tubu.) 39. Guaranitica (Licea I) 39. qiiroccpluila (Tilm.) 33'). gyrocephaluin (Didy.) 335. (ly)'Otmm (Phj's.) 307. Hariotii (Arcv.) 155. herbatica (Stem.) 87. heteratricJila (Aric.) 174. hians (Ailm.) 279. hians (Phys.) 296. Huolcri (i)iac.) 260. Hookeri (Dider.) 261. Hookeri (Lamp.) 261. hiunile (Didy.) 248. Lyalimi (Badh.) 319. hypnophiliun (Didy.) 224, hypiiophilum (Phys.) 314. inaurata (Badh.) 324. incamcda (Arcy.) 145. incarmitus (Lach.) 139. hiconspiciia (Trich.) 180. imitans (Phys. ) 292. inaequalis (Phys.) 300. iiiermis (Arcy.) 157. insignis (Arcy.) 148. intermedia (Trich.) 188. intricata (Arcy.) 144. intricata (Crib.) 59. iridescens (Lamp.) 101. iridei(/m (Lamp.) 95. irrecjnlaris (Badh.) 323. irregidarls (Ophi.) 132. irregularis (Peri.) 133. irregularis (Arcj\) 146. Jackii (Trich.) 188. Kalhreyeri (Trich.) 191. Kalrhhreiitieri (Phys.) 297. A'r'r.s/r//((Arcy.) 168. K!ck.rli (Trich.) 195. Kxrzii (Lepi.) 255. laciniatum (Dider.) 201, lateritia (Trich.) 176. laxa (Stem.) 79. laxa (Coma.) 79. leicorpa (Hemi.) 1G7. leicorpa (Arcy.) 167, leocarpoides (Corn.) 167. leocarpuidi's (Arcy.) 167. Leprieurii (Arcy.) 153. leptotrichum (Chon.) 244. leucuphaeiun (Phys.) 288. leptotrichum (Didy.) 243. leucucepliahini (Crat.) 267. leucophaeum (Phys.), o. flexuosuni, 287. leucophaeum (Phys. ) 288. leucupoda (Diac.) 259. leucopus (Phys.) 287. leihcosporum (Lamp.) 96. Leveillei (Phys.) 296. liceoides (Peri.) 118. Uceoides (Chon.) 215. licheniformis (Spum.) 292. lilacina (Badh.) 271. Ulacinnm (Crat.) 271. Liudheimeri (Licea) 42. Lindheimeri (Tubu ) 42. Lisicri (Lamp.) 97. Listeri (Didy.) 244. lividnm (Phys.) 304. lividum (Phys.) 2S4. lobatum (Didy.) 220. longa (Coma.) 83. lonya (Stem.) 83. longipes (Didy.) 236. Lorinseriana (j3.) 175. lucidum (Dider.) 204. lucidum (Chon.) 204. luteolvm (Phys.) 309. luteo-valvc (Phys.) 310. Lyallii (Chon.) 201. lycoperdi (Crib.) 104. hjcoperdon (Retic.) 93. lycopodii (Lamp.) 104. macrocarpa (Badh.) 317. macrocarpa (Cril).) 56. macrocarpon (Phys.) 317. nuicrosperma (Stem.) 76. inacrospermum (Didy.) 228. macro.'ipxiriim (Enter.) 45. magna (Badh.) 319. magnum (JJicty.) 319. viaxima (Brei'.) 91. Index. 365 maxima (Stem.) 74. mdnnopeziza (Arcy.) 162. melanopeziza (Hemi.) 162. mdanospom (Badh.) 325. inelleum (Didy.) 278, 279. melUnm (Didy.) 276, 278. metallioa (Corn.) 130. mdallira (Proto.) 128. mutallica (Tricli.) 128. inetallicum (Phys.) 130. Michel a (Chon.) 204. Michelii (Dider.) 205. 'microcarpa (Badh.) 325. microcarpa (Crib.) 63. microcarpa (Peri.) 117. nucrocarpou (Didy.) 227. micrcnirpon (Didy.) 226. mirroaphala (Ortli.) 109. microscop'ica (Crib.) 62. niicrosperma (Licea) 40. minima (Crib.) 59. minima (Tubu.) 36. minima (Licea) 36. minnna (Trich.) 182. m.iiuituna (Crat.) 272. minuta (Tilm.) 296. minuta (Tilm.) 279. minutissima (Crib.) 66. minutulum (Olig.) 171. minutnm (Lamp.) 100. mimUnm (Ecliin.) 109. minutum (Lyco.) 125. minutnm (Crat.) 264. minntum (Crat.) 273. mirabile (Heter.) 60. mirahilis (Crib.) 60. mirabilis (Phys.) 334. Mortjaiu (Stem.) 86. muscicolnin (Phys.) 312. miisconiiii (Ener.) 106. muscorum (Phys.) 307. mutabilc (Chon.) 212. mutal)ile (Crat.) 269. mntabile (Crat.) 272. mntubUis (Tilm.) 329. nana (Trich.) 181. nefroideiim (Phvs.) 285. neglect nm (Didy.) 231. neglectum (Didy.) 224. nigrescens (Lamp.) 96. nigrescens(Lamp.) 102. nigripes (a.) 227. nitcns (Badh.) 324. nitem (Ophi.) 13,3. nitens (Peri.) 133. nifeus (dig.) 170. nitens (Olig.) 173. nilens (Trich.) 179. nitidissima (Tubu.) 30. nitidum (Lyco.) 122. nimnm (Chon.) 206. nivevm (Chon.) 207. nodvlom (Badh.) 322. i,o/ c'.s (Chon.) 214. plujsarioides (Lamp.) 103. physarioides (Stem.) 96. :^6G A Mo)iofj)'ap]i of tlie Myxogastres. physarioiihs (Didy.) 233. picemn (Pbys.) 313. pirifannis (Cril).) 55. platypus (Anaricus) 239. 2)latiipvs (Didy.) 239. polyacdron (Phys.) 316. po'ymorjihum (Phys.) 283. polymorpliuni (Phys.) 251. pomiformis (Arcy.) 151. praecoM (Didy.) 223. pro'ximella (Trich.) 180. proximum (Didy.) 238. pruinosiim (Didy.) 288. pseudaecidium (Peri.) 119. ■psittacumm (Phys.) 274. pnlcheUa (Stem.) 86. pulchella (Coma.) 86. jjulcherrimnm (Phys.) 293. pttlcherripes (Phys.) 315. puuicm (Arcy.) 142. punicea (Arcy.) 149. pii/rpiirascens (Tricli.) 177. purjnorea (Crib.) 57. pusilla (Hemi.) 168. pusilla (Arcy.) 168. pusilla (Hemi.) 168. pusilla (Trich.) 195. pusilla (Proto.) 44. pusillum (Proto.) 44. 2)nsUlum (Proto.) 43. pusillum (Didy.) 238. pyriforme (Crat.) 264. quercina (Peri.) 115. Eaciborsckii (Arcy.) 156. radlatnm (Didy.) 229. radiatmn (Chon.) 200, 202. Eavenelii (Didy.) 281. Ravendil (Phys.) 281. Eeaderi (Phys.) 282. reniformis (Trich.) 184. rnuformis (Tilm.) 336. reticulata (Licea) 253. reticulata (Licea) 133. reticulata (Ophi.) 133. reticulata (Peri.) 133. r('//c^'/((^M/H (Chon.) 216. reticnlutuiii (Didy.) 253. rrticuldtxtn (Lepi.) 252. reflcidatimi (Phys.) 314. robvda (Lamp.) 99. roseiim (Phys.) 294. liostufinsliii (Lach.) 138, Rodajitiskii (Peri.) 117. RostafinsUi (Phys.) 301. Rostnijiii (Enter.) 45. Rozeana (Relic.) 47. ii''ovo( /(Hi, (Enter.) 47. rubiformis (Arcy.) 158. rnbiijinosa (Badh.) 270. ntbigiuosum (Crat.) 270. rnhiginosvin (Phys.) 302. riibifjinosHm (Scyphium) 270. nifa (Crib.) 63. rufibasis (Phys.) 296. rvjibasis (Phys.) 279. rufipes (jS ) 227. rnfo-cinnamomeiim (Lyco.) 125. rngidosum (Clat.) 51. Saccardianum (Lamp.) 101. Sanndersii (Chon.) 209. Sauteri (Chon.) 217. Santeri (Lamp.) 100. scabra (Trich.) 192. Schimperi (Lamp.) 99. Schroeteri (Phys. ) 280. Schnnuicheri (Phys.) 275. Schumacher! (Phys.) 335. Schumacheri (Phys.) 278. Schweinitzii (Corn.) 52. Scluocmitzii (Phys.) 311. scrobiculatum (Didy.) .301. scrobicidatum (Phys.) 300. scyphoides (Phj's.) 282. serpida (Arcy.) 164. serpula (Arcy.) 136. seipuJa (Didy.) 234. scpvla (Ophi.) 135. serpula (Hemi.) 165. simile (Phys.) 286. simdis (Arcy.) 156. simplex (Chon.) 217. smiulans (Chon.) 209. simulans (Enter.) 47. sinapinicm (Didy.) 246. sinuosum (Phys.) 305. spadiceum (O.str.) 52. speciosa (Tubu.) 38. spleHdens (Crib.) 62. splendens (Stem.) 74. siAendens (Diac.) 261. spermoides (Licea) 37. spermoides (Tnlm.) 37. spumarioidea (Licea) 42. spumarioidea (Tubu.) 42. spumarioidcs (Didy.) 232. Imle^r. 367 spumarioides (Cliit,((,i (Tubu.) 38. slipitata (Spor.) 219. stipitatuiu (y.) 277. strarainifonuis (Crib.) 335. striatum (Phys.) 243. stricta (Arcy.) 151. stromateum (Chon.) 233. snlir„rsi>;f,>s,( (Stem.) 80. subdictvospermniu (Clion.) 208. iinhfii.^ra (Trich.) 183. sub.4b,lM,sum (Phys.) 300. s>illif,, -11 1 urn {Chon.) 211. subucncinn. (Lamp.) 95. subroseum (Didy.) 312. siibsessilis (Diac.) 262. Suksdorfii (Coma.) 76. Suk^dor/li (Stem.) 76. svlphicrea {Tvich.) 186. sidphnreum (Phys.) 311. superba (Tricli.) 194. Tatrica (Crib.j 62. tenella (Crib.) 58. tenerrima (Stem.) 81. 'est ICewman otroiTuj.litii West.Nevonan chr.lith i'iV3t,Nevmia.n cnr lith West.NevwmKui chr lith W-sl f.ewnian I'h' %3 ^ ^" West, Neviman chr. lith ^. ♦ *a "^ ^yj^J.a "" ' "- Hg'-^i^ / i>*^€';\- . - v.r-'w .' ' ''■' ' „ * , r ^^^^^^^^^^^R ^A ^^ V- ^^^^^^^^^^^^^^K ^ < Is ^^^^^^% '4'M~ ;^'' 1 "5 ■Wf" i w